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  • †Tuzoia sinensis P'an 1957




    Kingdom: Animalia
    Phylum: Arthropoda Latreille 1829
    Class: incertae sedis
    Order: Tuzoida Simonetta and Delle Cave, 1975
    Family: Tuzoiidae Raymond, 1935
    Genus: Tuzoia
    Species: Tuzoia sinensis
    Author Citation P'an 1957

    Geological Time Scale

    Eon: Phanerozoic
    Era: Paleozoic
    Period: Cambrian
    Sub Period: None
    Epoch: Series 2 Epoch
    International Age: Stage 4


    Wulongqing Formation


    Acquired by: Purchase/Trade


    Length: 5 cm


    Chenggong District, Kunming Town


    Taxonomy from Vannier et al. (2007, p. 458).

    Vannier et al. 2006 assume that Tuzoia and the also Cambrian genus Isoxys are possibly representatives of the class Thylacocephala.

    Revised diagnosis for the genus Tuzoia Walcott, 1912 in Vannier et al. (2007, pp. 458–459): “Large bivalved arthropod (valve length up to approx. 180 mm; Chlupác and Kordule, 2002). Carapace thin, nonmineralized, and relatively flexible, folded dorsally (no true articulated hinge) into two hemispherical valves. Valve outline amplete or slightly preplete/postplete (L:H ratio between 1.3 and 1.6). Straight or slightly convex dorsal margin with or without spines. Anterior and posterior cardinal processes (acp and pcp, respectively) pointed in most species. Acp usually more prominent than pcp and with underlying notch. Posterior margin typically with a mid-posterior spine (mps) and a posteroventral spine (pvs). Angle between dorsal margin and pvs between 20° and 40°, most frequently 30°. In some species, additional small marginal spines (sms) and crenulation, especially ventrally, their length decreasing towards the front. Well-marked to vestigial lateral ridge along the entire length of the valves, usually at mid-distance between dorsal and ventral margin on a direct line with mps. In some species, lateral ridge armed with a series of 7–10 thin and broad-based, probably flimsy, spines (spines projecting almost perpendicular to the sagittal plane of the carapace). Valve surface with well-marked reticulate pattern (size of polygonal fossae decreasing towards lateral ridge, cardinal areas, and along margins). A pair of large, stalked lateral eyes and short antennae protruding through anterior notch. Eye stalks at least three times longer than the eye itself. Gut with possible digestive glands.”

    General morphology of Tuzoia Walcott, 1912 from Vannier et al. (2007, p. 446):



    Identified by oilshale using Luo et al., 2006.


    P’an, K., (1957): On the discovery of Homopoda from South China. Acta Palaeontologica Sinica 5, 523–526.

    LUO, H.L., FU, X.P., HU, S.X., LI, Y., CHEN, L.Z., YOU, T. & LIU, Q., (2006): New bivalved arthropods from the early Cambrian Guanshan fauna in the Kunming and Wuding area. Acta Palaeontologica Sinica 45, 460–472.

    Vannier, J., Caron, J.-B., Yuan J.-L., Briggs, D. E. G., Collins, D., Zhao Y.-L., Zhu M.-Y. (2007): Tuzoia: Morphology and Lifestyle of a Large Bivalved Arthropod of the Cambrian Seas. Journal of Paleontology 81 (3), 445–471.

    Zhao, J., Li, Yujing, Selden, Paul A. and Cong, Peiyun (2020): New occurrence of the Guanshan Lagerstatte (Cambrian Series 2, Stage 4) in the Kunming area, Yunnan, southwest China, with records of new taxa. ALCHERINGA: AN AUSTRALASIAN JOURNAL OF PALAEONTOLOGY, VOL. 44, NO. 3, 343–355 DOI: 10.1080/03115518.2020.1781257.

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