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  1. Taxonomy from Wang et al. 2009. Daouhugounectes primitivus is a predaceous water beetle grub with raptorial forelegs and natatorial mid- and hindlegs. Diagnosis from Wang et al. 2009, p. 653." Head transverse, narrowing anterior of the base; nasale with rounded lateral lobes. Mandible with retinaculum near its middle. Prothoracic tergite somewhat longer than meso- and metathoracic tergites. Legs relatively short; mid- and hindleg distinctly longer than foreleg, their femora, tibiae and tarsi subequal in length, tibae slightly dilated at apex, tarsi roundly dilated anteriorly. Forelegs with sparse strong setae; meso- and metafemora with sparse thin hairs at posterior margin, tibiae and tarsi with short oblique setae and long think hairs along both sides. Abdomen with nine segments, seven of them transverse; sclerites in tergal position; sclerite VIII almost circular, curved ventromedially. Small subquadratic segment IX present between urogomphi. The closing apparatus of the spiracles on segment VIII weakly developed; trachea with helical thickening weak (compared to that of Coptoclava). Urogomphi approximately 2.5 times as long as tergite VIII." Line drawing from Wang et al, 2009 (scale bar 5 mm): Reference: B. Wang, A. G. Ponomarenko, and H.-Ch. Zhang (2009): A new coptoclavid larva (Coleoptera: Adephaga: Dytiscoidea) from the Middle Jurassic of China, and its phylogenetic implication. Paleontological Journal 43(6):652-659.
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