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  1. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slab. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin, 2001 reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian); this opinion is now widely accepted. Taxonomy according to Fossilworks.org. Emended diagnosis for Choristopsychidae according to Qiao et al., 2013, p. 93: "Forewing broad oval or subtriangular, field between C and ScP comparatively broad; ScP well developed and forked twice, forming three long branches; RA unforked, one crossvein between ScP and RA and between RA and RP; RP and MA both with two branches; MP with five branches, and the MP4+5 forking basal to the MP2+3 forking; MP and CuA merged at the base; CuA strongly bent at its mid point; an oblique crossvein between CuA and CuP; a curved crossvein between the midpoint of CuA and MP5; CuP, 1A and 2A almost parallel. Hind wing, similar in shape to the forewing but slightly smaller, ScP short, forked twice, the second bifurcation coalesces with RA for a short distance; RP and MA both with two branches; MP with five branches, the stem of MP4+5 forked earlier than that of forewing, and with a crossvein to CuA; CuA almost straight. Head, oviform with big and oval compound eyes; antennae long and filiform; small chewing mouthpart. Thorax: prothorax smaller than mesothorax and metathorax. Legs: long and slender, all legs nearly of the same shape, but hind legs longer than fore legs and mid legs, and femora wider than tibia, and tibia longer than femora. Abdomen slender, tapering apically, about eleven segments and the female terminal segment with cercus." Choristopsyche asticta, line drawing of the holotype from Qiao et al., 2013, p. 100. Identified by Prof. A. Rasnitsyn (Russian Academy of Sciences) as Choristopsyche cf. asticta Qiao et al., 2013. Reference: Gao K Q, Shubin N. (2001): Late Jurassic salamanders from Northern China. Nature, 410: 574–577. Martynov, A. V. (1937): Liassic insects from Shurab and Kisyl-Kiya, Part I, Various orders except Blattodea and Coleoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7: 1-178. Qiao, X., Shih, C. K., Petrulevičius, J. F., and Ren, D. (2013): Fossils from the Middle Jurassic of China shed light on morphology of Choristopsychidae (Insecta, Mecoptera). ZooKeys 318: 91-111. link
  2. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slab. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian), this opinion is now widely accepted. Taxonomy according to Wei et al., 2012. Line drawing from Wei et al., 2012. Description according to Wei et al., 2012: “Small-sized, body length about 10.4–10.6 mm (with head), width 2.8–3.1 mm; head small, significantly elongated (length/width= 1.4–1.6 mm/1.3–1.4 mm), antennal socket conspicuous at sides, mouthparts unclear; pronotum length 1.6–1.9 mm, width 2.3–2.7 mm, elliptical, as wide as the body; abdomen 6–7 segments visible, terminal sternum rounded; long cerci has 14 segments and apex of cerci strongly curved inward and rounded in shape, forming a narrow gap at center (Fig 3A), segments of cerci joined together after the 8th segment. Forewings (Figs 1, 2, 3B): length range about 8.5–8.8 mm, width range about 2.6–2.9 mm; narrow, without coloration, with intercalaries and wing venation simple, with 30–32 veins at margin; costal area wide (1/3 width of the wing); Sc simple, curved upward, longer than clavus; R strongly curved like waves and with 9–14 branches, reaching the anterior wing margin; M slightly curved and with 5–7 branches, most posterior branch of M reaching wing apex; CuA almost straight to posterior wing margin and with 5–8 branches; CuP strongly curved and simple; clavus short, less than a third of the wing’s length; A simple, arc bending and with about 4 veins.” Identified by oilshale using Wei et al., 2012. References: Wei D. D., Liang J. H. and Ren D. (2012) A new species of Fuziidae (Insecta, Blattida) from the Inner Mongolia, China. ZooKeys 217: 53–61. Gao, K. -Q. and Shubin, N. H. (2012) Late Jurassic salamandroid from western Liaoning, China. Proceedings of the National Academy of Sciences. 109 (15): 5767–72.
  3. Part and counterpart of the same fossil. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slabs. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin 2012 reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian), this opinion is now widely accepted. Taxonomy according Fossilworks.org. Line drawing from Liang et al., 2009, p.20. Description according to Liang et al., 2009, p. 19: “Large Species (forewing length/width: 21.1–26.0mm/6.5–8.0mm; hind wing length/width: 21.0–24.0mm/7.0–8.3mm; head length/width: 3–4.6mm/2–2.5mm; pronotum length/width: 5–6.2mm/4.5mm–5.8mm), with body strongly sclerotised. Head base wide and coloured, eyes located basally, partially covered by pronotum; ocelli invisible. Pronotum vaulted, slightly elongate with dark coloration at center and at margins. Veins dark, intercalaries and cross-veins distinct. Forewing with expanded venation. Area between anterior wing margin and Sc very long and narrow, Sc branched; R with undifferentiated Rs, does not reach apex; CuP slightly curved. Anal veins with tertiary branches, the base of anal region colored. Diagonal fold present. Hind wing with simple Sc; R differentiated into R1 and well-developed Rs; M reduced to few, usually 2 branches. CuA basal most branches strong, CuP simple, straight; anal lobe with fan-like pleating, A1 simple. Reticulations present in CuA-CuP space, joined with intercalaries. Body wide. Abdominal segments long, with parallel margins. Cerci multi-segmented, the last three segments longest. Ovipositor outer valves internalized.“ Identified by oilshale using Liang et al., 2009. References: Liang, J. H., Vršanský, P., and Ren, D. (2009): A new Jurassic carnivorous cockroach (Insecta, Blattaria, Raphidiomimidae) from the Inner Mongolia in China. Zootaxa 1974:17-30. Gao, K. -Q. and Shubin, N. H. (2012): Late Jurassic salamandroid from western Liaoning, China. Proceedings of the National Academy of Sciences. 109 (15): 5767–72.
  4. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin reported an Argon-argon dating age of 164+- 4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian), this oppinion is now widely accepted. Part and counterpart of the same fossil. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slabs. Drawing of the holotype by Liang et al. 2012 Taxonomy according Fossilworks.org. Differential diagnosis (genus) according to Liang et al, 2012: "The new genus can be distinguished from other representatives of the family by the richly branched Sc. The new genus additionally differs from Liadoblattina Handlirsch, 1906 (Vršanský and Ansorge 2007) from Early Jurassic of Germany and England in having a larger body and reticulations present in the CuA-CuP area. Graciliblatta is also different from Raphidiomima and Cameloblatta in the elongated pronotum (the length-to-width ratio of the pronotum of the new genus is 1.5, those of Raphidiomima and Cameloblatta 1.2), and distinctive divided eyes located basally, which are undivided in the other genera, which originate from the Late Jurassic of Kazakhstan. The head and pronotum of Graciliblatta are extremely long and narrower than in Fortiblatta (Liang et al. 2009); vein R of the forewing is more arcuate and R1 has secondary branching not found in Fortiblatta, which is from the same locality and age as Graciliblatta. In addition, the body is smaller (length/width of head in the new genus is 2.25 vs. 1.9 in Fortiblatta; the pronotal length/width of the new genus is 1.5 vs. 1.2 in Fortiblatta). Coloration in the tip of wings is absent, but Graciliblatta has two dark stripes on pronotum, unlike Fortiblatta." Description (species) according to Liang et al., 2012: "Head and pronotum elongated. Forewing long and narrow. Sc richly branched, with 7–8 branches; R not reaching the tip of margin, slightly curved and basally with dark coloration. CuA and CuP slightly curved. Anal veins with tertiary branches. Diagonal fold present. Hind wing with simple Sc; R1 and Rs differentiated, and R1 secondarily branched." Prognathous and sharp mandibles of the species suggest it was a carnivore. Identified by oilshale. Reference: LIANG, J. H., HUANG, W. L. & REN, D. (2012) Graciliblatta bella gen. et sp. n. - A rare carnivorous cockroach (Insecta, Blattida, Raphidiomimidae) from the Middle Jurassic. Zootaxa 3449: 62–68. LIANG, J. H., SHIH, C. K. & REN D. (2018). New Jurassic predatory cockroaches (Blattaria: Raphidiomimidae) from Daohugou, China and Karatau, Kazakhstan. Alcheringa 42:101-109.
  5. Taxonomy from Wang et al. 2009. Daouhugounectes primitivus is a predaceous water beetle grub with raptorial forelegs and natatorial mid- and hindlegs. Diagnosis from Wang et al. 2009, p. 653." Head transverse, narrowing anterior of the base; nasale with rounded lateral lobes. Mandible with retinaculum near its middle. Prothoracic tergite somewhat longer than meso- and metathoracic tergites. Legs relatively short; mid- and hindleg distinctly longer than foreleg, their femora, tibiae and tarsi subequal in length, tibae slightly dilated at apex, tarsi roundly dilated anteriorly. Forelegs with sparse strong setae; meso- and metafemora with sparse thin hairs at posterior margin, tibiae and tarsi with short oblique setae and long think hairs along both sides. Abdomen with nine segments, seven of them transverse; sclerites in tergal position; sclerite VIII almost circular, curved ventromedially. Small subquadratic segment IX present between urogomphi. The closing apparatus of the spiracles on segment VIII weakly developed; trachea with helical thickening weak (compared to that of Coptoclava). Urogomphi approximately 2.5 times as long as tergite VIII." Line drawing from Wang et al, 2009 (scale bar 5 mm): Reference: B. Wang, A. G. Ponomarenko, and H.-Ch. Zhang (2009): A new coptoclavid larva (Coleoptera: Adephaga: Dytiscoidea) from the Middle Jurassic of China, and its phylogenetic implication. Paleontological Journal 43(6):652-659.
  6. Northern Sharks

    Bug ID request

    While this isn't the type of "bug" I normally collect, this one appealed to me as soon as I saw it on the auction site. Can anyone in our masses give me any help with regards to identification? The seller, who is also a member on here BTW, speculated at the order Neuroptera, but that was followed by a (?). It is Jurassic in age, from the Daohugou lagerstatte of Inner Mongolia. Thanks in advance.
  7. oilshale

    Orthophlebia sp.

    Is either related or identical to Orthophlebia stigmosa (picture A) or O. nervulosa (picture D) Reference.: Xiao Qiao, Chungkun Shih & Ren (2012): Two new Middle Jurassic species of orthophlebiids (Insecta: Mecoptera) from Inner Mongolia, China. Alcheringa: An Australasian Journal of Palaeontology, DOI:10.1080/03115518.2012.671689
  8. From the album: Invertebrates

    Same story as yesterday: My colleague and I wanted to visit a customer near Nincheng in Nei Mongol, China when I saw a drywall made out of grey flag stones. Looked very promising. I could pursuade my colleague (who was driving the car) to take a rest.... Unfortunately I could not tear down the wall - the owner of the drywall didn't like my idea and opposed. I guess the shale is from Daohugou region. Mecoptera non det. Female Possibly belonging to the Panorpidae Jiulongshan Formation Middle Jurassic Ningcheng Nei Mongol China
  9. References: J. H. Liang, P. Vrsansky, and D. Ren (2012). Variability and symmetry of a Jurassic nocturnal predatory cockroach (Blattida: Raphidiomimidae). Revista Mexicana de Ciencias Geológicas 29:411-421
  10. From the album: Invertebrates

    Divocina noci Liang, Vršanský & Ren, 2012 Middle Jurassic Daohugou Nei Mongol PRC
  11. oilshale

    Insect non det. (Cicada?)

    From the album: Invertebrates

    Insect non det. Middle Jurassic Daohugou Nei Mongol PRC
  12. oilshale

    Insect non det.

    From the album: Invertebrates

    Insect non det. Middle Jurassic Daohugou Nei Mongol PRC
  13. oilshale

    Insect non det.

    From the album: Invertebrates

    Insect non det Upper Jurassic Daohugou Nei Mongol China
  14. oilshale

    Blattodea non det.

    From the album: Invertebrates

    Blattodea non det. Middle Jurassic Daohugou Nei Mongol China Length 1cm
  15. oilshale

    Insect non det.

    From the album: Invertebrates

    Insect non det. Middle Jurassic Daohugou Nei Mongol PRC
  16. Taxonomy from Gao & Shubin 2003. Identified by Prof Wang Yuan, Institute of Vertebrate Paleontology and Paleoanthropology, Beijing with the following remarks: 'The phalangeal formula for Chunerpeton is variable. This specimen falls in the range as I observed hundreds of specimens of this genus and species. One problem is that, as I noted in the last email, Gao and Shubin (2012) named Beiyanerpeton jianpingensis but it is actually a Chunerpeton by my judgement. The salamander you send to me should be from the same region, but a different quarry as "Beiyanerpeton".' Diagnosis from Gao & Shubin 2003, p. 425: "Chunerpeton tianyiensis shares with living cryptobranchoids derived characters including: presacral vertebrae bearing unicapitate ribs; reduction in the number of rib-bearing anterior caudal vertebrae reduced to two or three. Chunerpeton tianyiensis shares with cryptobranchids derived characters such as: nasal much narrower than interorbital width; nasal–prefrontal contact absent; frontals extend anteriorly to lateral border of nasal; lachrymal absent; anterolateral process of parietal extends along lateral border of frontal; internal carotid foramina penetrate palatal surface of parasphenoid. Chunerpeton tianyiensis differs from extant cryptobranchids in lacking midline contact of dorsal processes of premaxillae; frontal–maxillary contact absent; absence of contact between anterolateral process of parietal and prefrontal; vomers without posterior extension; retention of palatal fenestra between vomers; presence of distinct medial process of pterygoid; pterygoid– parasphenoid contact absent; basibranchial II ossified and trident-shaped; first three pairs of ribs with spatulate distal end; phalangeal formula of 2-2-3-(3/4)-3 in pes." Line drawing from Gao & Shubin 2003, p. 425: The white mass next to the mouth seems to be either a regurgitate or a coprolite. References: K.-Q. Gao and N. H. Shubin. 2003. Earliest known crown-group salamanders. Nature 422:424-428. Yuan Wang, Liping & Susan E. Evans (2014) Polydactyly and other limb abnormalities in the Jurassic salamander Chunerpeton from China.Palaeobio Palaeoenv DOI 10.1007/s12549-015-0219-7.
  17. Taxonomy from Liu et al. 2006. Diagnosis (Liu et al. 2006, p. 550): "Comparing with A. conspecta (Sinitshenkova, 1992),A. yehae sp. nov. exposits more morphological characters of the genus, and is distinct from A. conspecta in the forewings: RS forks almost at the level of the SC tip, rs-m entering the RS before the RS fork and touching M at the M fork, the shorter c-sc and the longer wing." Line drawing from Liu et al. 2006, p. 551: Identified by oilshale References: Liu et al. (2011): Pronemouridae fam. nov. (Insecta: Plecoptera), the stem group of Nemouridae and Notonemouridae, from the Middle Jurassic of Inner Mongolia, China. Palaeontology, Vol. 54, Part 4, 2011, pp. 923–933. Liu, Ren, Sinitshenkova & Shih (2006) A new Middle Jurassic stonefly from Daohugou, Inner Mongolia, China (Insecta: Plecoptera)  Annales Zoologici 56(3):549-554. Liu, Shih & Ren. 2010. In Ren, Shih, Zhao & Y Zhao. Silent Stories - Insect Fossil Treasures from Dinosaur era of the Northeastern China 65
  18. From the album: Invertebrates

    My colleague and I wanted to visit a customer near Nincheng in Nei Mongol, China when I saw a drywall made out of grey flag stones. Looked very promising. I could pursuade my colleague (who was driving the car) to take a rest.... Unfortunately I could not tear down the wall - the owner of the drywall didn't like my idea and opposed. I guess the shale came from Daohugou. Colorifuzia agenora Wei, Liang & Ren, 2013 Middle to Late Jurassic Jiulongshan Formation Daohugou Nei Mongol China
  19. References: Wei D. D., Liang J. H. & Ren D. (2013). A new fossil genus of Fuziidae (Insecta, Blattida) from the Middle Jurassic of Jiulongshan Formation, China. Geodiversitas 35 (2): 335-343. http://dx.doi.org/10.5252/g2013n2a3.
  20. oilshale

    Mecoptera non det.

    From the album: Invertebrates

    Mecoptera non det. Possibly an Orthophlebiidae according to Xiaodan Lin and Weiting Zhang (thanks to both!) Middle Jurassic Jiulongshan Formation Chifeng Province Nei Mongol PR China
  21. oilshale

    Cicadomorpha non det.

    From the album: Invertebrates

    Cicadomorpha non det. Anthoscytina sp? Middle Jurassic Jiulongshan Formation Chifeng Province Nei Mongol PR China
  22. From the album: Invertebrates

    Pseudopolycentropus daohugouensis ZHANG, 2005 Middle Jurassic Jiulongshan Formation Daohugou Nei Mongol China Lit.: Grimaldi, D.A., Junfeng, Z., Fraser, N.C., Rasnitsyn, A., 2005. Revision of the bizarre Mesozoic scorpionflies in the Pseudopolycentropodidae (Mecopteroidea). Insect Syst. Evol. 36, 443–458. Ren, D., Shih, C., Labandeira, C.C., 2010. New Jurassic pseudopolycentropodids from China (Insecta: Mecoptera). Acta Geol. Sinica 84, 22–30. Shih CK, Yang XG, Labandeira CC, Ren D (2011) A new long-proboscid genus of Pseudopolycentropodidae (Mecoptera) from the Middle Jurassic of China and its plant-host specializations. In: Shcherbakov DE, Engel MS, Sharkey MJ (Eds) Advances in the Systematics of Fossil and Modern Insects: Honouring Alexandr Rasnitsyn. ZooKeys 130: 281–297. doi: 10.3897/zookeys.130.1641
  23. References: Grimaldi, D.A., Junfeng, Z., Fraser, N.C., Rasnitsyn, A., 2005. Revision of the bizarre Mesozoic scorpionflies in the Pseudopolycentropodidae (Mecopteroidea). Insect Syst. Evol. 36, 443–458. Ren, D., Shih, C., Labandeira, C.C., 2010. New Jurassic pseudopolycentropodids from China (Insecta: Mecoptera). Acta Geol. Sinica 84, 22–30. Shih CK, Yang XG, Labandeira CC, Ren D (2011) A new long-proboscid genus of Pseudopolycentropodidae (Mecoptera) from the Middle Jurassic of China and its plant-host specializations. In: Shcherbakov DE, Engel MS, Sharkey MJ (Eds) Advances in the Systematics of Fossil and Modern Insects: Honouring Alexandr Rasnitsyn. ZooKeys 130: 281–297. doi: 10.3897/zookeys.130.1641
  24. References: Liu et al. (2011): Pronemouridae fam. nov. (Insecta: Plecoptera), the stem group of Nemouridae and Notonemouridae, from the Middle Jurassic of Inner Mongolia, China. Palaeontology, Vol. 54, Part 4, 2011, pp. 923–933.
  25. oilshale

    Insect non det.

    From the album: Invertebrates

    Insect non det. Middle Jurassic Daohugou Nei Mongol China
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