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  1. From the album: Invertebrates

    Jiania crebra Wang, Szwedo & Zhang, 2012 Hemiptera Froghopper Middle Jurassic Daohugou Inner Mongolia PRC
  2. From the album: Invertebrates

    Neimenggucossus normalis Wang, Zhang & Fang, 2007 Middle Jurassic Daohugou Inner Mongolia PRC
  3. From the album: Invertebrates

    Jiania crebra Wang, Szwedo & Zhang, 2012 Middle Jurassic Daohugou Inner Mongolia PRC
  4. Taxonomy from Fossilworks.com. Diagnosis from Liu, Y.s., Sinitshenkova, N.D. & Ren, D., 2009, p. 185: "Adults. Medium-sized stoneflies. Head large, antennae long, prothorax narrow. Wings at rest extending considerably beyond apex of abdomen. In forewings, c–r absent; Rs bifurcating, usually forked at level of Sc apex. M branching slightly distad of Rs fork; m–cu always connecting base of MP with CuA. Crossveins between M and CuA and between CuA and CuP numerous. CuP straight, entering wing margin approximately at level of M fork. Hindwings with four longitudinal veins in anal area. Legs long and thin. First tarsomere long, almost twice as long as second; second tarsomere shorter than third. Cerci short, single-segmented." Line drawing from Liu et al. 2009, p. 189: References: N. D. Sinitshenkova. (1987). Istopicheskoe razvitie vesiyanok. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 221:1-142. Liu, Y.s., Sinitshenkova, N.D. & Ren, D. (2009). A revision of the Jurassic Stonefly Genera Dobbertiniopteryx Ansorge and Karanemoura Sinitshenkova (Insecta: Plecoptera), with the description of new species from the Daohugou locality, China. Paleontol. J. 43, 183–190. https://doi.org/10.1134/S0031030109020099
  5. From the album: Invertebrates

    Xyelotoma macroclada Gao, Ren & Shih, 2009 Middle Jurassic Callovian - Oxfordian Jiulongshan Formation Daohugou Inner Mongolia PR China
  6. oilshale

    Daohugocorixa vulcanica Zhang, 2010

    From the album: Invertebrates

    Daohugocorixa vulcanica Zhang, 2010 Jurassic Callovian - Oxfordian Jiulongshan Formation Daohugou Inner Mongolia China
  7. Taxonomy from Chen et al. 2019. Diagnosis from Wang et al., p. 1233: "Tegmen c. 3.5 times as long as wide, with apex widely rounded; basal cell c. 0.17 times as long as tegmen length, closed with anastomosis; common stalk ScR + M very short; branch ScRA c. 1.5 times as long as stem ScR; stem CuA at base distinctly convex, forking basad of claval apex; stigmal cell narrow, c. half as wide as radial cell." Line drawing from Wang et al. 2012, p. 1227: References: WANG, B., SZWEDO, J. & ZHANG, H. (2012). NEW JURASSIC CERCOPOIDEA FROM CHINA AND THEIR EVOLUTIONARY SIGNIFICANCE (INSECTA: HEMIPTERA) Palaeontology, Vol. 55, Part 6, 2012, pp. 1223–1243] Chen, J., Wang, B., Zheng, Y., Jiang, H., Jiang, T., Zhang, J. Q., An, B. Z. and Zhang, H. C. (2019). New fossil data and phylogenetic inferences shed light on the morphological disparity of Mesozoic Sinoalidae (Hemiptera, Cicadomorpha). Organisms Diversity & Evolution 19:287-302 [M. Clapham/M. Clapham]
  8. oilshale

    Parakseneura sp.

    Taxonomy from Fossilworks.com. Diagnosis for the genus from Yang et al. 2012, p. 4: 'Large neuropterans (forewing 50–75 mm long) with the following character states: labial palpi stout, relatively short; antennae stout, filiform, apparently much shorter than forewing length; two tibial spurs straight, shorter than basitarsus; claws big, strongly curved; in both wing, humeral veinlet well-developed, strongly recurrent, branched; presumed ScA short, fused with ScP within humeral area; membrane covered with dense, long hairs; RA (or ScP+RA) entering margin well before wing apex; subcostal crossveins numerous; radial crossveins irregularly spaced, not forming gradate series; in the forewing, MP, CuA, CuP dichotomously branched; presumed AA1+2 very short (found in Parakseneura gen. nov.); AA3+4, AP1+2, AP3+4 deeply forked; in hind wing, presumed AA1+2 very short (found in Pseudorapisma gen. nov.); proximal half of hind wings considerably wider than distal.' Determined by Dr. V. Makarkin, Academy of Sciences Vladivostok, Russia, as Paraksneura sp. It might be P. albadelta. Line drawing from Yang et al., 2012, p. 12: References: Yang Q, Makarkin VN, Winterton SL, Khramov AV, Ren D. (2012) A Remarkable New Family of Jurassic Insects (Neuroptera) with Primitive Wing Venation and Its Phylogenetic Position in Neuropterida. PLoS ONE 7(9): e44762. doi:10.1371/journal.pone.0044762.
  9. Taxonomy from Fossilworks.org. Etymology: Derived from the Latin word formos- (beautiful) and Bittacus (a recent genus of Bittacidae). The species is named macularis after the presence of many maculae on wing. Diagnosis for the species †Formosibittacus macularis from Li et al. 2008, p. 42: "Sc very long, terminating at about three-fifth of the length of wing; sc-r at about two times its length before end of Sc; dark zones along cross-veins and darkened apex in wing membrane." Line drawing of body with wings, left forewing and left hindwing from Li et al., p. 41: References: Y. L. Li, D. Ren, and C. K. Shih. 2008. Two Middle Jurassic hanging-flies (Insecta: Mecoptera: Bittacidae) from northeast China. Zootaxa 1929: 38-46.
  10. Taxonomy from Mindat.org. Alternative name Netropanorpodes decorosus Sun, Ren & Shih 2007. Revised diagnosis from Cao et al. (2015), p. 9: "In forewings, Sc forks with three branches; R2+3 forks after the crossvein r1–r2+3; Cu1+M forks before the crossvein cu1–cu2. In hind wings, R2+3 forks before the crossvein r1–r2." Line drawing of the left forewing from Cao et al. 2015, p. 8: References: NAN LIU, YUNYUN ZHAO & REN (2010) Two new fossil species of Itaphlebia (Mecoptera: Nannochoristidae) from Jiulongshan Formation, Inner Mongolia, China. Zootaxa 2420: 37–45. CAO, Y., SHIH, C., BASHKUEV, A. & REN, D., XX.XX.XXXX. Revision and two new species of Itaphlebia (Nannochoristidae: Mecoptera) from the Middle Jurassic of Inner Mongolia, China. Alcheringa 40, XX–XX. ISSN 0311-5518. Cao Y, Lin X, Shih C, Ren D (2022) Two new species of Itaphlebia (Insecta, Mecoptera, Nannochoristidae) from the late Middle Jurassic of China. ZooKeys 1108: 175–188. https://doi.org/10.3897/zookeys.1108.85378
  11. Taxonomy from Fossilworks.org. Male scorpionfly. Species name reflects the characteristic pattern marking of the wings. Diagnosis from Soszyńska-Maj et al. 2019, p. 9: "Species differs from remaining congeneric by combination of the following characters: numerous oval-shaped small regular dark spots on the membrane all situated between veins spread evenly throughout the whole wings, more transparent areas than dark, Rs forks slightly beyond Mb forks in forewing, Rs1+2 almost twice as long and Rs3+4, basal part of M4b in forewing and M4 in hindwing strongly oblique and situated with cross-vein m-cu on one line." Line drawing from Soszyńska-Maj et al. 2019, p. 13 : References: Agnieszka Soszyńska-Maj, Wiesław Krzemiński, Katarzyna Kopeć, Yizi Cao, Ren, Wiesław Krzemiński & Katarzyna Kopeć (2019): New Middle Jurassic fossils shed light on the relationship of recent Panorpoidea (Insecta, Mecoptera), Historical Biology, DOI: 10.1080/08912963.2018.1564747
  12. Taxonomy from Fossilworks.org. Etymology. From “ellip- ”, which means elliptic and genus Ragio, referring to the elliptic wings. Gender: masculine. Diagnosis from Han et al. 2019, p.154: “Flagellum with 10 flagellemeres; proboscis long, labium fleshy, labella small. Wings elliptic and wide; R2+3 sinuate at the middle, and sharply up-curved distally; crossvein r-m intersecting the upper margin of d cell at basal one third (1/3); four medial veins present, bM3 and dM3 straight; anal cell closed before wing margin. Midtibiae with 1 spur.“ Line drawings from Han et al., 2019, p. 157 (scale bar equals 0.5mm): References: Han YE, Cai Y, Ren D, Wang Y. (2019). A new fossil snipe fly with long proboscis from the Middle Jurassic of Inner Mongolia, China (Diptera: Rhagionidae). Zootaxa. 30;4691(2):zootaxa.4691.2.4. Zhang, K., Yang, D. & Ren, D. (2008). A new genus and species of Middle Jurassic rhagionids from China (Diptera, Rhagionidae). Biologia, 63(1), 113-116. https://doi.org/10.2478/s11756-008-0012-4 Zhang, J. (2013). Snipe flies (Diptera: Rhagionidae) from the Daohugou Formation (Jurassic), Inner Mongolia, and the systematic position of related records in China. Palaeontology, 56, 1, 217–228. Zhang, K., Li, J., Yang, D., & Ren, D. (2009). A new species of Archirhagio Rohdendorf, 1938 from the Middle Jurassic of Inner Mongolia of China (Diptera: Archisargidae). Zootaxa, 1984, 61-65.
  13. oilshale

    Mesosciophila eucalla Zhang 2007

    Taxonomy from Fossilworks.org. Emended diagnosis for the genus from Zhang 2007, p. 298: "Medium-sized mesosciophilid gnats. Male body (including legs) covered with long, dense pubescence. Eyes large. Maxillary palps five-segmented, longer than head length. Antennae filiform, 16-segmented, with scapes and pedicels quadrate, flagellomeres cylindrical. Mesonotum convex. Scutellum clearly projecting. Venationally, Sc1 ending distad to level of Rs origin, Sc2 situated clearly basad to Rs origin; bRs longer than r-m; R1 slightly curved; both R1 and R4+5 divergent terminally; Rs furcated distad to fork of M1+2; R2+3 oblique; cell r moderately large, one-quarter to one-fifth of length of wing; stem of M not developed. Halteres light, with pubescence not visible. Femora, tibiae and first two tarsomeres with one or two rows of short setae." Line drawings from Zhang 2007, p. 300 (scale bars represent 1mm): A: Male Gnat C: Wing References: Junfeng Zhang (2007). New mesosciophilid gnats (Insecta: Diptera: Mesosciophilidae) in the Daohugou biota of Inner Mongolia, China, Cretaceous Research, Volume 28, Issue 2, Pages 297-301, ISSN 0195-6671, https://doi.org/10.1016/j.cretres.2006.05.007.
  14. oilshale

    Itaphlebia laeta Liu et al. 2010

    Taxonomy from Mindat.org. Diagnosis from Liu et al. 2010: "Costal area narrow. In forewing, Sc simple, terminating at C near pterostigma; M with 4 branches; Cu 1 and M forked before the cross-vein cu 1 -cu 2." Description from Liu et al. 2010: "Antenna filiform, incompletely preserved. Femur shorter than tibia. Tibial spurs not present. Setae irregularly arranged. Costal area narrow, one cross-vein c-sc between C and Sc. Sc simple, ending at C and extending to pterostigmal area. One short cross-vein sc 2 -r 1 before pterostigma. Rs arised at the same level to cross-vein csc. Rs with 4 branches. Length of R 2 two-thirds that of stem of R 2 + 3. Cross-vein r 1 -r 2 at level of pterostigma. Cross-vein r 3 -r 4 at level of cross-vein r 1 -r 2. Cross-vein r 5 -m 1 + 2 approximately at middle of wing length. Crossvein r 5 -m 1 proximal to cross-vein r 4 -r 5. One cross-vein r 4 -r 5 dividing cell R 4 into 2 cells. Conspicuous thyridium at fork of M, which has 4 branches. M 3 + 4 divided beyond the fork of M 1 + 2; cross-vein m 1 + 2 -m 3 between M 1 + 2 and M 3; cross-vein m-cu not straight, slightly S-shaped. Cu 1 coalesced with M for a short distance and separated from M before cross-vein cu 1 -cu 2. One cross-vein a 1 -a 2 connected 1 A and 2 A. 3 A absent. Female abdomen with 11 segments, segments 9–11 smaller than segment 8. One pair of cerci located at apex of abdomen. Right cercus with 3 segments. Third segment of left cercus not preserved. Basal segments fused with abdomen segment 11. Body length 7.8 mm, forewing length 8.2 mm, width 2.6 mm." Line drawing of the left forewing from Liu et al. 2010: Reference: NAN LIU, YUNYUN ZHAO & REN (2010) Two new fossil species of Itaphlebia (Mecoptera: Nannochoristidae) from Jiulongshan Formation, Inner Mongolia, China. Zootaxa 2420: 37–45.
  15. Taxonomy according to Hao et al 2009. Hao et al. 2009, p223: “Revised diagnosis: Body length: 6-13 mm: wing length 8.2-15.6 mm Sc ending at wing margin approximately the same as R4+5 forking into R4 and R5; R1 long, r-r at one -third of its length before the end of R1; R2+3 shorter than R3; R4+5 short, R4+5 fork six times longer than dR4+5. M1 smoothly curved, m-m joining close to M1+2 bifurcations. M3+4 with a little bend at m-m. CuA slightly sigmoid beyond m-cu; A1 strongly curved to wing margin in two-thirds of its length.” Wing line drawing from Hao et al. 2009, p.225: Identified by oilshale using Hao et al. 2009. References: Ren, D. & Krzemiński, W. 2002. Eoptychopteridae (Diptera) from the Middle Jurassic of China. Ann. Zoologica 52 (2): 207-210. Hao J., Ren, D. and Chungkun S. (2009) New Fossils of Eoptychopteridae (Diptera) from the Middle Jurassic of Northeast China Acta Geologica Sinica 83(2):222 – 228 DOI: 10.1111/j.1755-6724.2009.00022.x Liu, L., Shih, C. and Ren, D. (2012) Two new species of Ptychopteridae and Trichoceridae from the Middle Jurassic of northeastern China (Insecta: Diptera: Nematocera). Zootaxa 3501: 55-62, DOI: 10.5281/zenodo.282475
  16. oilshale

    Osmylidae indet.

    There are additional unidentified conchostraca (clam shrimps, arthropods) on the slab. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin, 2001 reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian); this opinion is now widely accepted. Taxonomy according to Fossilworks.org. Diagnosis for the family OSMYLIDAE Leach, 1815 according to Winterton et al., 2019 p. 13. “Adult head with dorsal tentorial arms weakly developed; ocelli usually present, medial ocellus sometimes reduced (e.g., Paryphosmylus) or all ocelli completely absent (e.g., Gumillinae); palpimaculae absent; antennae filiform; wings with basal subcostal veinlet simple in costal area, sometimes forked but never recurrent; single basal crossvein (sc-r) between Sc and R, rarely more than one (e.g., Porismus, Archaeosmylidia); Sc and RA fused apically and joining costal margin before wing apex in both wings; single branch of RP, subsequently pectinately branched (5 or more branches); FW radial branches frequently curved posteriorly in apical half of wing, sometimes sinuous; FW medial vein fork variably placed along wing, HW medial vein fork near wing base; FW CuP pectinately (rarely dichotomously) branched; cubital field relatively large in both wings; nygmata typically present; trichosors present in both wings (rarely reduced to apical margin); end twigging along wing margin present, sometimes extensive and multi-layered; jugal lobe present; female gonocoxite 9 (gx 9) (i.e., gonopophysis lateralis) elongate and with terminal stylus; female with a pair of sclerotised spermathecae; male genitalia with well-developed arched gonarcus with paired entoprocesses fused laterally; mediuncus curved; parameres present or absent, sometimes fused into single arched sclerite; larva (Fig. 1) with straight, highly elongate jaws; gular-like sclerite absent; seven Malphigian tubules, of which five arge incorporated into the cryptonephridium; eversible claws present on paired prolegs on last abdominal segment.” Identified by Alex Khramov (Borissiak Paleontological Institute of the Russian Academy of Sciences, Moscow, Russia): “Osmylidae, but definitely not Ponomarenkius or any other Kempyninae. Which genus or subfamily I cannot tell you - too fragmentary preservation and, moreover, there are many undescribed osmylids in Daohugou, so probably your specimen is one of them.” References: Gao K Q, Shubin N. (2001): Late Jurassic salamanders from Northern China. Nature, 410: 574–577. Winterton, Shaun L., Martins, Caleb C., Makarkin, V., Camacho, Adrian A., & Wang, Yongjie (2019): Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera. Zootaxa 4581 (1): 001–099. https://doi.org/10.11646/zootaxa.4581.1.1 .
  17. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slab. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin, 2001 reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian); this opinion is now widely accepted. Taxonomy according to Fossilworks.org. Emended diagnosis for Choristopsychidae according to Qiao et al., 2013, p. 93: "Forewing broad oval or subtriangular, field between C and ScP comparatively broad; ScP well developed and forked twice, forming three long branches; RA unforked, one crossvein between ScP and RA and between RA and RP; RP and MA both with two branches; MP with five branches, and the MP4+5 forking basal to the MP2+3 forking; MP and CuA merged at the base; CuA strongly bent at its mid point; an oblique crossvein between CuA and CuP; a curved crossvein between the midpoint of CuA and MP5; CuP, 1A and 2A almost parallel. Hind wing, similar in shape to the forewing but slightly smaller, ScP short, forked twice, the second bifurcation coalesces with RA for a short distance; RP and MA both with two branches; MP with five branches, the stem of MP4+5 forked earlier than that of forewing, and with a crossvein to CuA; CuA almost straight. Head, oviform with big and oval compound eyes; antennae long and filiform; small chewing mouthpart. Thorax: prothorax smaller than mesothorax and metathorax. Legs: long and slender, all legs nearly of the same shape, but hind legs longer than fore legs and mid legs, and femora wider than tibia, and tibia longer than femora. Abdomen slender, tapering apically, about eleven segments and the female terminal segment with cercus." Choristopsyche asticta, line drawing of the holotype from Qiao et al., 2013, p. 100. Identified by Prof. A. Rasnitsyn (Russian Academy of Sciences) as Choristopsyche cf. asticta Qiao et al., 2013. Reference: Gao K Q, Shubin N. (2001): Late Jurassic salamanders from Northern China. Nature, 410: 574–577. Martynov, A. V. (1937): Liassic insects from Shurab and Kisyl-Kiya, Part I, Various orders except Blattodea and Coleoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7: 1-178. Qiao, X., Shih, C. K., Petrulevičius, J. F., and Ren, D. (2013): Fossils from the Middle Jurassic of China shed light on morphology of Choristopsychidae (Insecta, Mecoptera). ZooKeys 318: 91-111. link
  18. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slab. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian), this opinion is now widely accepted. Taxonomy according to Wei et al., 2012. Line drawing from Wei et al., 2012. Description according to Wei et al., 2012: “Small-sized, body length about 10.4–10.6 mm (with head), width 2.8–3.1 mm; head small, significantly elongated (length/width= 1.4–1.6 mm/1.3–1.4 mm), antennal socket conspicuous at sides, mouthparts unclear; pronotum length 1.6–1.9 mm, width 2.3–2.7 mm, elliptical, as wide as the body; abdomen 6–7 segments visible, terminal sternum rounded; long cerci has 14 segments and apex of cerci strongly curved inward and rounded in shape, forming a narrow gap at center (Fig 3A), segments of cerci joined together after the 8th segment. Forewings (Figs 1, 2, 3B): length range about 8.5–8.8 mm, width range about 2.6–2.9 mm; narrow, without coloration, with intercalaries and wing venation simple, with 30–32 veins at margin; costal area wide (1/3 width of the wing); Sc simple, curved upward, longer than clavus; R strongly curved like waves and with 9–14 branches, reaching the anterior wing margin; M slightly curved and with 5–7 branches, most posterior branch of M reaching wing apex; CuA almost straight to posterior wing margin and with 5–8 branches; CuP strongly curved and simple; clavus short, less than a third of the wing’s length; A simple, arc bending and with about 4 veins.” Identified by oilshale using Wei et al., 2012. References: Wei D. D., Liang J. H. and Ren D. (2012) A new species of Fuziidae (Insecta, Blattida) from the Inner Mongolia, China. ZooKeys 217: 53–61. Gao, K. -Q. and Shubin, N. H. (2012) Late Jurassic salamandroid from western Liaoning, China. Proceedings of the National Academy of Sciences. 109 (15): 5767–72.
  19. Part and counterpart of the same fossil. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slabs. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin 2012 reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian), this opinion is now widely accepted. Taxonomy according Fossilworks.org. Line drawing from Liang et al., 2009, p.20. Description according to Liang et al., 2009, p. 19: “Large Species (forewing length/width: 21.1–26.0mm/6.5–8.0mm; hind wing length/width: 21.0–24.0mm/7.0–8.3mm; head length/width: 3–4.6mm/2–2.5mm; pronotum length/width: 5–6.2mm/4.5mm–5.8mm), with body strongly sclerotised. Head base wide and coloured, eyes located basally, partially covered by pronotum; ocelli invisible. Pronotum vaulted, slightly elongate with dark coloration at center and at margins. Veins dark, intercalaries and cross-veins distinct. Forewing with expanded venation. Area between anterior wing margin and Sc very long and narrow, Sc branched; R with undifferentiated Rs, does not reach apex; CuP slightly curved. Anal veins with tertiary branches, the base of anal region colored. Diagonal fold present. Hind wing with simple Sc; R differentiated into R1 and well-developed Rs; M reduced to few, usually 2 branches. CuA basal most branches strong, CuP simple, straight; anal lobe with fan-like pleating, A1 simple. Reticulations present in CuA-CuP space, joined with intercalaries. Body wide. Abdominal segments long, with parallel margins. Cerci multi-segmented, the last three segments longest. Ovipositor outer valves internalized.“ Identified by oilshale using Liang et al., 2009. References: Liang, J. H., Vršanský, P., and Ren, D. (2009): A new Jurassic carnivorous cockroach (Insecta, Blattaria, Raphidiomimidae) from the Inner Mongolia in China. Zootaxa 1974:17-30. Gao, K. -Q. and Shubin, N. H. (2012): Late Jurassic salamandroid from western Liaoning, China. Proceedings of the National Academy of Sciences. 109 (15): 5767–72.
  20. A sensation: First fossil evidence of a hybrid of Odonata and Arachnida from the Middle Jurassic of Daohugou, Inner Mongolia. A talented artist has painted one pair of legs too much by mistake. Was unfortunately sold to a trusting collector.
  21. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin reported an Argon-argon dating age of 164+- 4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian), this oppinion is now widely accepted. Part and counterpart of the same fossil. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slabs. Drawing of the holotype by Liang et al. 2012 Taxonomy according Fossilworks.org. Differential diagnosis (genus) according to Liang et al, 2012: "The new genus can be distinguished from other representatives of the family by the richly branched Sc. The new genus additionally differs from Liadoblattina Handlirsch, 1906 (Vršanský and Ansorge 2007) from Early Jurassic of Germany and England in having a larger body and reticulations present in the CuA-CuP area. Graciliblatta is also different from Raphidiomima and Cameloblatta in the elongated pronotum (the length-to-width ratio of the pronotum of the new genus is 1.5, those of Raphidiomima and Cameloblatta 1.2), and distinctive divided eyes located basally, which are undivided in the other genera, which originate from the Late Jurassic of Kazakhstan. The head and pronotum of Graciliblatta are extremely long and narrower than in Fortiblatta (Liang et al. 2009); vein R of the forewing is more arcuate and R1 has secondary branching not found in Fortiblatta, which is from the same locality and age as Graciliblatta. In addition, the body is smaller (length/width of head in the new genus is 2.25 vs. 1.9 in Fortiblatta; the pronotal length/width of the new genus is 1.5 vs. 1.2 in Fortiblatta). Coloration in the tip of wings is absent, but Graciliblatta has two dark stripes on pronotum, unlike Fortiblatta." Description (species) according to Liang et al., 2012: "Head and pronotum elongated. Forewing long and narrow. Sc richly branched, with 7–8 branches; R not reaching the tip of margin, slightly curved and basally with dark coloration. CuA and CuP slightly curved. Anal veins with tertiary branches. Diagonal fold present. Hind wing with simple Sc; R1 and Rs differentiated, and R1 secondarily branched." Prognathous and sharp mandibles of the species suggest it was a carnivore. Identified by oilshale. Reference: LIANG, J. H., HUANG, W. L. & REN, D. (2012) Graciliblatta bella gen. et sp. n. - A rare carnivorous cockroach (Insecta, Blattida, Raphidiomimidae) from the Middle Jurassic. Zootaxa 3449: 62–68. LIANG, J. H., SHIH, C. K. & REN D. (2018). New Jurassic predatory cockroaches (Blattaria: Raphidiomimidae) from Daohugou, China and Karatau, Kazakhstan. Alcheringa 42:101-109.
  22. References: J. H. Liang, P. Vrsansky, and D. Ren (2012). Variability and symmetry of a Jurassic nocturnal predatory cockroach (Blattida: Raphidiomimidae). Revista Mexicana de Ciencias Geológicas 29:411-421
  23. Echinoid

    Epidexipteryx hui

    A friend bought me some Copic artists' markers recently, so I decided to draw Epidexipteryx hui. It is a strange little creature from the Daohugu beds of the Jurassic of China. It is based off an illustration by Fabio Pastori in The Bumper Book of Dinosaurs. Apologies for the lines - for some reason I draw better on lined paper Inspiration:
  24. From the album: Invertebrates

    Divocina noci Liang, Vršanský & Ren, 2012 Middle Jurassic Daohugou Nei Mongol PRC
  25. oilshale

    Insect non det. (Cicada?)

    From the album: Invertebrates

    Insect non det. Middle Jurassic Daohugou Nei Mongol PRC
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