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  1. Taxonomy according to fossilworks.org. Diagnosis for the genus from Bannikov 2008, p. 79: "Body depth is 13-14 times in SL; head length 2.9 times in SL; teeth moderate, conical; 56 vertebrae; 2 epurals; epaxial hypurals unfused; distance between dorsal fins exceeds distance between second dorsal and caudal fin; 12 to 13 interdorsal bones between dorsal fins; a spine and 7 or 8 soft rays in second dorsal fin; a spine and 9 or 10 soft rays in anal fin; second dorsal fin originates not less than 2 vertebrae behind anal-fin origin; pectoral fins short, with 13 or 14 rays; 6-rayed pelvic fins moderate, situated about 4 vertebrae anterior to first dorsal-fin origin; pectoral to pelvic fin distance less than pelvic to anal fin distance." Line drawing from Bannikov 2008, p. 83: Identified by oilshale using Bannikov 2008. Confirmed by A. F. Bannikov, Paleontological Institute of the Russian Academy of Sciences, Moscow. References: Agassiz, L. (1835) Recherches Sur Les Poissons Fossiles. Tome IV (livr. 4). Imprimerie de Petitpierre, Neuchatel 33-52. Bannikov A.F., (2008e). Revision of the atheriniform fish genera Rhamphognathus Agassiz and Mesogaster Agassiz (Teleostei) from the Eocene of Bolca, northern Italy. Miscellanea Paleontologica n. 9. Studi e Ricerche sui Giacimenti Terziari di Bolca, 12: 77-97. Carnevale, G., Bannikov, A. F., Marramá, G., Tyler, J. C. and Zorzin, R. (2014): The Pesciara-Monte Postale Fossil-Lagerstätte: 2. Fishes and other vertebrates. Rendiconti della Società Paleontologica Italiana 4:37-63. Bannikov. A. F. (2014): The systematic composition of the Eocene actinopterygian fish fauna from Monte Bolca, northern Italy, as known to date. Studi e ricerche sui giacimenti terziari di Bolca, XV - Miscellanea paleontologica 12:23-34.
  2. Taxonomy according to Marramà & Carnevale 2015. Diagnosis for the genus Bolcaichthys according to Marramà & Carnevale 2015: “Small- to medium-sized clupeid (largest specimen measuring 101.1 mm SL) with elongated and tapered body, sardine-like; † Bolcaichthys, gen. nov., is unique among clupeids in having the following combination of characters: head length approximately one third to one quarter SL; skull roof with 10–14 frontoparietal striae; mouth terminal; two supramaxillae, anterior small and rod-like, and posterior paddle-shaped; teeth absents in jaws and palate; complete series of abdominal keeled scutes (10–11 prepelvic and 10 or 11 postpelvic) with ascending arms; dorsal scutes absent; fewer branchiostegal rays (5 or 6), supraneurals (8), vertebrae (40–42), and pleural ribs (20–22) compared with other genera of clupeids; pleural ribs–preural vertebrae ratio ranging from 0.48 to 0.55; three epurals; deeply forked caudal fin with 19 principal caudal-fin rays; dorsal fin small, inserting at about mid-length of the body with 15 or 16 rays; about 15 or 16 anal-fin rays; about 14–18 pectoral-fin rays; pelvic-fin origin slightly in front of or behind the posterior end of the dorsal fin; eight pelvic-fin rays.” Line drawing of Bolcaichthys catopygopterus (Woodward, 1901) from Marramà and Carnevale, 2015: Identified by oilshale using Marramà & Carnevale, 2015. References: Agassiz, L. (1833–1844). Recherches sur les Poissons Fossiles. Petitpierre, Neuchâtel, 1420 pp. Woodward, A. S. (1901): Catalogue of Fossil Fishes in the British Museum (Natural History), 4: Containing the Actinopterygian Teleostomi of the Suborders Isospondyli (in part), Ostariophysi, Apodes, Percesoces, Hemibranchii, Acanthopterygii and Anacanthini. Taylor and Francis, London, 636 pp. Marramà, G. & Carnevale, G. (2015): The Eocene sardine †Bolcaichthys catopygopterus (Woodward, 1901) from Monte Bolca, Italy: osteology, taxonomy, and paleobiology, Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2015.1014490
  3. Taxonomy according Fossilworks.org. Diagnosis according to Blot 1978, p. 146: "Tête contenue un peu moins de six fois dans la longueur totale. Hauteur du corps contenue de quinze à dix-neuf fois dans la même longueur. Les deux branches de cleithra de longueurs équivalentes. Côtes ventrales présentes. Osselets intermusculaires bien développés. Caudale constituée par neuf rayons s'articulant sur les hypuraux. Coefficient de concentration de la nageoire dorsale: 2,50, celui de l'anale étant aux environs de 2,85." This translates: “Head contained a little less than six times in the total length. Height of the body contained from fifteen to nineteen times in the same length. Both cleithra branches of equivalent lengths. Ventral ribs present. Intermuscular bones well developed. Caudal constituted by nine rays articulating on the hypurals. Coefficient of concentration of the dorsal fin: 2.50, that of the anal being around 2.85.” Line drawing of Bolcyrus formosissimus from Blot 1978, p. 149. Identified by oilshale using Blot 1978. References: Blot, J. (1976) Les anguilliformes fossiles du Monte Bolca. 2e Congres Europeen des Ichtyologistes Europeens, Paris, 1976, Revue Trav. Inst. Pech. Marit., Nantes, 40 (3&4) 509-511, 1 tabl. Blot, J. (1978): Les apodes fossiles du Monte Bolca. Studi e Ricerche sui Giacimenti Terziari di Bolca 3:(1): 1-260, 120 fig, 21 tabl. 39pl. Blot, J. (1984): Les Apodes fossiles du Monte Bolca. 2. Actinopterygii : Ordre des Apodes (Anguilliformes): Famille des Paranguillidae Blot 1980. Museo civico di storia naturale di Verona, 1984, p. 62-238, 24 p. di tav. G. Carnevale, A. F. Bannikov, G. Marramá, J. C. Tyler, and R. Zorzin (2014): The Pesciara-Monte Postale Fossil-Lagerstätte: 2. Fishes and other vertebrates. Rendiconti della Società Paleontologica Italiana 4:37-63.
  4. References: SECRETAN S (1975) Les crustacés de Monte Bolca. Studi e Ricerche sui Giacimenti Terziari di Bolca 2. Museo Civico di Storia Naturale di Verona: 315-388.
  5. Hello everybody can somebody help me identifying my fossil fish (4cm) I am pretty convinced that it concerns the exellia velifer from monte bolca looking forward to Your replies regards Rienk
  6. oilshale

    Carangopsis dorsalis

    From the album: Vertebrates

    Carangopsis dorsalis Middle Eocene Monte Bolca Verona Italy length 10cm
  7. References: Blainville H.M.D. (de), (1818) Poissons fossiles. Chap. VII. Des Ichthyolites du bassin de la Méditerranée. a. Des Ichthyolithes du Monte Bolca, ou Vestena Nuova dans le Veronais in Nouveau dictionnaire d'histoire naturelle, appliquée aux Arts, à l'Agriculture, à l'Economie rurale et domestique, à la Médecine, etc. Nouvelle Edition presqu'entièrement refondue et considérablement augmentée, t. 27, p. 334-361. Woodward A.S., (1901) Catalogue of the fossil fishes in the British Museum, t. 4. Blot J., (1980) La faune ichthyologique des gisements du Monte Bolca (Province de Vérone, Italie). Bulletin du Muséum national d'Histoire naturelle: 339-396, fasc. 4, sér. 4, t. 2, part. section C. G. Carnevale, A. F. Bannikov, G. Marramá, J. C. Tyler, and R. Zorzin (2014) The Pesciara-Monte Postale Fossil-Lagerstätte: 2. Fishes and other vertebrates. Rendiconti della Società Paleontologica Italiana 4:37-63. A. F. Bannikov (2014) The systematic composition of the Eocene actinopterygian fish fauna from Monte Bolca, northern Italy, as known to date. Studi e ricerche sui giacimenti terziari di Bolca, XV - Miscellanea paleontologica 12:23-34
  8. oilshale

    Ductor vestenae Volta 1796

    Taxonomy from Fossilworks.org Line drawing from Blot 1969, Tome 2, Planche B: Identified by oilshale using Blot 1969. References: Blot, J. (1969): Les poissons fossiles du Monte Bolca : classes jusqu'ici dans les familles des Carangidae, Menidae, Ephippidae, Scatophagidae / Jacques Blot. - Verona : Museo Civico di Storia Naturale di Verona, 1969. – Tome 1 and 2. G. Carnevale, A. F. Bannikov, G. Marramá, J. C. Tyler, and R. Zorzin (2014) The Pesciara-Monte Postale Fossil-Lagerstätte: 2. Fishes and other vertebrates. Rendiconti della Società Paleontologica Italiana 4:37-63
  9. oilshale

    Ductor vestenae VOLTA, 1796

    From the album: Vertebrates

    Ductor vestenae VOLTA, 1796 Eocene Ypresian Monte Bolca near Verona Italy
  10. oilshale

    Rhamphosus rastrum (VOLTA, 1796)

    Alternative spelling: Ramphosus References: L. Agassiz (1842) Recherches Sur Les Poissons Fossiles. Tome IV (livr. 14). Imprimerie de Petitpierre, Neuchâtel 205-291. G. Carnevale, A. F. Bannikov, G. Marramá, J. C. Tyler, and R. Zorzin (2014) The Pesciara-Monte Postale Fossil-Lagerstätte: 2. Fishes and other vertebrates. Rendiconti della Società Paleontologica Italiana 4:37-63 [M. Uhen/M. Uhen/M. Uhen]
  11. A few weeks ago I was looking for a new case for my iPhone. So I browsed around a Chinese internet shop. I came across cases with fossil pictures on the back: Icaronycteris from the Green River Formation, a Seymouria from the Permian, Aipichthys velifer from the Cretaceous of Lebanon, Archaeopteryx from Solnhofen, a fossil turtle (Green River?) and finally a Sparnodus vulgaris from the Eocene of Monte Bolca, Italy. Funny only that this Sparnodus vulgaris seemed so familiar to me! No wonder - the original is in my collection and hangs on the wall in the corridor. The Chinese company searched for some templates in the internet and found them at TFF. They probably downloaded the picture here in the forum. Now I have an IPhone housing with the image of a fish from my own collection. Thomas
  12. A bottom dweller, similar to the recent Gobius. References: Bannikov A.F. (2004) Eocottidae, a new family of perciform fishes (Teleostei) from the Eocene of northern Italy (Bolca) // Studi ric. giacim. terz. Bolca. Verona. 2004. V. X. p. 17-35. Bannikov A.F. (2006) Bassanichthys, a new replacement generic name for the Eocene Bassania Bannikov, 2004 (Teleostei, Perciformes)// Paleontological Journal, Vol. 40, Issue 3, p. 340.
  13. oilshale

    Milananguilla lehmani Blot, 1975

    References: Blot, J. (1976) Les anguilliformes fossiles du Monte Bolca. 2e Congres Europeen des Ichtyologistes Europeens, Paris, 1976, Revue Trav. Inst. Pech. Marit., Nantes, 40 (3&4) 509-511, 1 tabl. Blot, J. (1978) Les apodes fossiles du Monte Bolca. Studi e Ricerche sui Giacimenti Terziari di Bolca, Verona 3 (1) 1-260, 120 fig, 21 tabl. 39pl. Blot, J. (1984) Les Apodes fossiles du Monte Bolca. 2. Actinopterygii : Ordre des Apodes (Anguilliformes): Famille des Paranguillidae Blot 1980. Museo civico di storia naturale di Verona, 1984, p. 62-238, 24 p. di tav.
  14. oilshale

    Ramphosus rastrum (Volta, 1796)

    From the album: Vertebrates

    Ramphosus rastrum (Volta, 1796) Middle Eocene Monte Bolca near Verona Italy
  15. oilshale

    cf. Prosolenostomus lessinii

    Taxonomy from Bannikov & Carnevale, 2017. Bannikov & Carnevale 2017, p. 319:" Its type species, †P. lessinii Blot, 1980, was described based on a single partially complete specimen in the collections of the Museo Civico di Storia Naturale, Verona (Blot, 1980, p. 365; pl. VI, fig. 3). Subsequently, an additional incomplete specimen housed in the Carnegie Museum of Natural History, Pittsburgh, was referred to †P. lessinii and figured by Frickhinger, 1991, p. 769); however, this specimen lacks the characteristic cranial horn emerging just above the orbit considered diagnostic by Blot (1980)." Photo of the holotype of Prosolenostomus lessinii in Bannikoy & Carnevale 2017, p. 320: This here is the third known specimen of Prosolenostomus. Like the specimen from the Carnegie Museum of Natural History, Pittsburgh, illustrated by Frickhinger 1991, p. 769, this Prosolenostomus has no cranial horn, and is therefore a new, as yet undescribed species. A. Bannikov wrote (personal communication, 13.12.2021) " the "cranial horn" is not a conservation artifact, but a genuine diagnostic feature. Therefore your specimen belongs to the till undescribed taxon. Overall, pipefishes are very rare in Monte Bolca." Diagnosis for the genus Prosolenostomus from Blot, 1980, p. 365 (translated from French by oilshale); "Very elongated body. Cuirass constituted by cruciform scutes, imbricated the ones in the others, delimiting more or less abundantly reticulated beaches. Very large branchial opening. Short head with very high rostral tube and terminal mouth; very strong mandibles. A rather high horn rises above the orbit. " Identified as cf. Prosolenostomus sp. by A. Bannikov (Borisyak Paleontological Institute, Russian Academy of Sciences). References: Blot J., (1980) La faune ichthyologique des gisements du Monte Bolca (province de Verone, Italie) : catalogue systematique presentant l´etat actuel des recherches concernant cette faune. Bulletin du Muséum national d´histoire naturelle. Section C, sciences de la terre, paléontologie, géologie, minéralogie, 4. serie, 2(4): 339-396. Frickhinger, K.-A (1991) Fossilien Atlas - Fische. Melle Mergus, ISBN 10: 388244018XISBN 13: 9783882440188 Alexandre F. BANNIKOV & Giorgio CARNEVALE (2017) Eocene ghost pipefishes (Teleostei, Solenostomidae) from Monte Bolca, Italy. Bollettino della Società Paleontologica Italiana, 56 (3), 2017, 319-331. Modena.
  16. oilshale

    Cyclopoma gigas Agassiz, 1833

    References: L. Agassiz (1833) Recherches Sur Les Poissons Fossiles. Tome IV (livr. 1). Imprimerie de Petitpierre, Neuchatel 17-32. G. Carnevale, A. F. Bannikov, G. Marramá, J. C. Tyler, and R. Zorzin (2014) The Pesciara-Monte Postale Fossil-Lagerstätte: 2. Fishes and other vertebrates. Rendiconti della Società Paleontologica Italiana 4:37-63. A. F. Bannikov (2014) The systematic composition of the Eocene actinopterygian fish fauna from Monte Bolca, northern Italy, as known to date. Studi e ricerche sui giacimenti terziari di Bolca, XV - Miscellanea paleontologica 12:23-34.
  17. Old name: Naseus rectifrons Agassiz, 1842 Taxonomy from Fossilworks.com. References: Agassiz, L. (1842) Recherches Sur Les Poissons Fossiles. Tome IV (livr. 14). Imprimerie de Petitpierre, Neuchâtel 205-291. Blot, J. and Tyler, J. C. (1990) New genera and species of fossil surgeon fishes and their relatives (Acanturoidei, Teleostei) from the Eocene of Monte Bolca, Italy, with application of the Blot formula to both fossil and recent forms. Studi e ricerche sui giacimenti terziari di Bolca 6:13-92. Carnevale, G., Bannikov, A. F., Marramá, G., Tyler, J. C. and Zorzin, R. (2014) The Pesciara-Monte Postale Fossil-Lagerstätte: 2. Fishes and other vertebrates. Rendiconti della Società Paleontologica Italiana 4:37-63. Bellwood D.R. (1996) The Eocene fishes of Monte Bolca: the earliest coral reef fish assemblage. Coral Reefs 15: 11-19.
  18. From the album: Vertebrates

    Eorandallius rectifrons (Agassiz, 1842) Middle Eocene Lutetian Monte Bolca near Verona Italy Length 10cm Lit.: J. Blot and J. C. Tyler. 1990. New genera and species of fossil surgeon fishes and their relatives (Acanturoidei, Teleostei) from the Eocene of Monte Bolca, Italy, with application of the Blot formula to both fossil and recent forms. Studi e ricerche sui giacimenti terziari di Bolca 6:13-92
  19. oilshale

    Sparnodus vulgaris Blainville, 1818

    Taxonomy from Fossilworks.com Diagnosis from Day 2002, p. 335: "Distinguished from other Eocene sparids by a robust jaw with large, conical teeth. Ethmoid dorsal margin is depressed directly anterior to the ethmoid±frontal suture. Deep-bodied form with a length to width ratio of 3:1. The supraneural formula appears to be 0/0 + 0/2 + 1/1; and the anterodorsal processes of the supraneurals overlap. Hypurals are separate. Caudal ®n is of low aspect ratio. Formula of the dorsal fin XII + 9; anal fin III + 9." Identified by oilshale using Day 2003. References: Bannikov A. F. (2006) Fishes from the Eocene of Bolca, northern Italy, previously classified in the Sparidae, Serranidae and Haemulidae (Perciformes). Geodiversitas 28 (2) : 249-275. Day J. J. (2002) Phylogenetic relationships of the Sparidae (Teleostei: Percoidei) and implications for convergent trophic evolution. Biological Journal of the Linnean Society 76: 269-301. Day J. J. (2003) Evolutionary relationships of the Sparidae (Teleostei: Percoidei): integrating fossil and Recent data. Transactions of the Royal Society of Edinburgh: Earth Sciences 93: 333-353. L. Agassiz (1839) Recherches Sur Les Poissons Fossiles. Tome IV (livr. 13). Imprimerie de Petitpierre, Neuchatel 109-204
  20. References: L. Agassiz (1839) Recherches Sur Les Poissons Fossiles. Tome IV (livr. 13). Imprimerie de Petitpierre, Neuchatel 109-204. Bannikov, A. F. ( 2014) The systematic composition of the Eocene actinopterygian fish fauna from Monte Bolca, northern Italy, as known to date. Studi e ricerche sui giacimenti terziari di Bolca, XV - Miscellanea paleontologica 12:23-34
  21. oilshale

    Eolates gracilis AGASSIZ, 1843

    Lates is a genus of freshwater and marine fish belonging to the order Perciformes. The lates species are now native to the Indian and western Pacific Oceans and to rivers and lakes in Africa. The lates species are generally carnivorous, feeding on invertebrates and other fish. The Nile perch, Lates niloticus, is one of the largest freshwater fish, it reaches a maximum length of nearly two metres (more than six feet), weighing up to 200 kg (440 lb).
  22. oilshale

    Sphyraena bolcensis AGASSIZ, 1844

    Recent Sphyraenea barracuda, own work of Laban712 (from Wikipedia): The genus Sphyraena (from Latin: "pike-like") belongs to the order Perciformes. Members of the genus Sphyraena, better known as Barracudas, are elongated fish, pike-like in appearance, with large pointed heads and jaws. The two dorsal fins are widely separated with the anterior fin having five spines, the posterior fin having only one spine and nine soft rays. The posterior dorsal fin is similar in size to the anal fin and is situated above it. Barracudas normally have an under bite with prominent sharp-edged fang-like teeth in sockets. The oldest known barracuda fossil found so far dates to the Eocene period. All members of the genus Sphyraena are voracious predators; they feed upon cephalopods and crustaceans but mainly prey on fishes. Young barracudas can be often seen in small schools. Adult barracudas are considered to be solitary when it comes to hunting. The great barracuda can swim up to 35 mph, yet it can move forward slowly with no apparent effort; thus increasing its stealth. Barracuda are found primarily in tropical and subtropical waters of the Atlantic, Indian, and Pacific oceans, but certain species such as the Great Barracuda lives in brackish water.
  23. Taxonomy from Fossilworks.org. The order Beryciformes, a poorly understood group, is represented in Monte Bolca with at least two species: Eoholocentrum and Berybolcensis, both from the subfamily Holocentrinae or squirrelfish (L. Sorbini, 1984). Both species seemed to be largely or entirely nocturnal and lived in deep marine waters; their eyes are amazingly large. References: Sorbini, L., 1979. Les Holocentridae du Monte Bolca. III. Berybolcensis leptacanthus (Agassiz). Studi e Richerche sui Giacimenti Terziari di Bolca 4, 19–35.
  24. oilshale

    Diodon tenuispinus AGASSIZ, 1833

    Diodon holocanthus, inflated (own work of Ibrahim lujaz from Rep. Of Maldives) Diodon nicthemerus (own work of user Springcold at da.wikipedia) Porcupinefish belong to the family Diodontidae within the Tetraodontiformes order and are also commonly called blowfish. They have the ability to swallow water or air and to inflate their body making it harder for predators to swallow them. When the fish inflates, sharp spines radiate outwards as a second defense mechanism. Some species are poisonous, having tetrodotoxin in their internal organs. Fugu is the Japanese word for pufferfish and is also a Japanese dish made out of the pufferfish meat. Because fugu is lethally poisonous if prepared incorrectly, fugu has become one of the most celebrated and notorious dishes in Japanese cuisine. Porcupinefish are medium to large sized fish and are found in shallow temperate and tropical seas worldwide. Monte Bolca is an important lagerstatte for Tetraodontiformes with Diodon tenuispinus as one of its oldest records. References: J. C. Tyler and F. Santini (2002) Review and reconstructions of the tetraodontiform fishes from the Eocene of Monte Bolca, Italy, with comments on related Tertiary taxa. Studi e Ricerche sui Giacimenti Terzari di Bolca. Museo Civico di Storia Naturale di Verona 9:47-119.
  25. Pesciarichthys baldwinae Sorbini & Tyler, 1998 was redescribed in 2012 by Bannikov & Tyler and the new genus Frigosorbinia established. Taxonomy from Fossilworks.org. Diagnosis from Bannikov and Tyler 1998, p. 27:" Body a deep ovoid in shape; head relatively small. Mouth small; teeth relatively small and lobate. Five branchiostegal rays. Vertebrae 22 (9 + 13); haemal spines of first two caudal vertebrae slender, oriented posteroventrally, and not in close proximity in their middle regions. Tenth caudal vertebra with elongate slender neural and haemal spines. One supraneural. Dorsal fin with eight spines and 23 soft rays. Both first dorsal-fin spine and first anal-fin spine protrude to exterior. Anal fin with three spines and 21 soft rays. Tenth interneural space with two basal pterygiophores; 11th space with one basal pterygiophore. First interhaemal space with two basal pterygiophores. Pelvic bone with relatively shallow subpelvic keel (12% of pelvic length). Scales very small over most of body and head. Each scale consists of a small basal plate with upright spinules. Larger rounded scale plates scattered posteriorly on body in region of caudal peduncle; these larger scales are densely spaced together and slightly more than twice the size of the small body scales elsewhere. Caudal peduncle with three moderate, fixed, keeled bony scutes." Line drawing from Sorbini & Tyler 1998, p. 25 : Identified by oilshale using Sorbini & Tyler 1998. References: Sorbini L. and Tyler, J. C. (1998) A new species of the Eocene surgeon fish genus Pesciarichthys from Monte Bolca, Italy, with comments on caudal penduncle armature and supraneurals in acanthurids. Studi e ricerche sui giacimenti terziari di Bolca 7:21-34. Bannikov. A. & Tyler, J.C. (2012): REDESCRIPTION OF THE EOCENE OF MONTE BOLCA, ITALY, SURGEON FISH PESCIARICHTHYS PUNCTATUS PERCIFORMES, ACANTHURIDAE, AND A NEW GENUS, FRIGOSORBINIA, FOR P. BALDWINAE. Studi e ricerche sui giacimenti terziari di bolca, XIV - Miscellanea Paleontologica, 11, 2012.
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