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  1. oilshale

    Prolyda elegantula Wang et al., 2016

    Taxonomy from Wang et al., 2016. Amended diagnosis from Wang et al., 2016, p. 73; "Head massive, circular or cube-like; mandibles curved, strong and sickle-like; pronotum short and wide; the first antennal flagellomere equal to head in length, but eight times as long as the second flagellomere; forewing pterostigma variable, completely sclerotized or partly sclerotized, or just membranous; M diverging from M+Cu at much larger angle than Cu; 1-RS proclival or somewhat vertical; angle between 1-M and RS+M almost 90°; 1cu-a distal to the middle of cell 1mcu or located at middle; 2r-rs almost in line with 2r-m; hind wing with 1r-m rather long, as long as or slightly shorter than 1-M." Line drawing from Wang et al. 2016, p. 77: References: Wang C, Shih C, Rasnitsyn AP, Wang M. (2016) Two new species of Prolyda from the Middle Jurassic of China (Hymenoptera, Pamphilioidea). Zookeys. 2016 Feb 26;(569):71-80. doi: 10.3897/zookeys.569.7249. PMID: 27110151; PMCID: PMC4829680.
  2. oilshale

    Osmylopsychopsidae non det.

    From Peng et al. 2015, p. 2: "The systematics of Mesozoic psychopsoids is very complicated and confused. This group of Neuroptera includes taxa with broad and multi-veined wings that are currently classified in five families, i.e. Osmylopsychopidae, Brongniartiellidae, Psychopsidae, Kalligrammatidae and Aetheogrammatidae (Makarkin et al. 2013). Kalligrammatidae and Aetheogrammatidae are easily distinguished from other psychopsoids, whereas the Mesozoic members of the three other families (i.e. Osmylopsychopidae, Brongniartiellidae and Psychopsidae) hardly differ from each other.". Determined in Mai 2023 by Dr. Vladimir N. Makarkin, Institute of Biology and Soil Sciences, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok, Russia: "This is a species of Osmylopsychopidae, possibly new." References: Makarkin, V. N., Yang, Q., Shi, C. F. & Ren, D. (2013). The presence of the recurrent veinlet in the Middle Jurassic Nymphidae (Neuroptera) from China: a unique condition in Myrmeleontoidea. ZooKeys, 325, 120. Peng, Y., Makarkin, V. N. and Ren, D. (2015). Diverse new Middle Jurassic Osmylopsychopidae (Neuroptera) from China shed light on the classification of psychopsoids. Journal of Systematic Palaeontology, p. 1-35. http://dx.doi.org/10.1080/14772019.2015.1042080
  3. From the album: Invertebrates

    Jiania crebra Wang, Szwedo & Zhang, 2012 Hemiptera Froghopper Middle Jurassic Daohugou Inner Mongolia PRC
  4. From the album: Invertebrates

    Neimenggucossus normalis Wang, Zhang & Fang, 2007 Middle Jurassic Daohugou Inner Mongolia PRC
  5. From the album: Invertebrates

    Jiania crebra Wang, Szwedo & Zhang, 2012 Middle Jurassic Daohugou Inner Mongolia PRC
  6. Taxonomy from Fossilworks.com. Diagnosis from Liu, Y.s., Sinitshenkova, N.D. & Ren, D., 2009, p. 185: "Adults. Medium-sized stoneflies. Head large, antennae long, prothorax narrow. Wings at rest extending considerably beyond apex of abdomen. In forewings, c–r absent; Rs bifurcating, usually forked at level of Sc apex. M branching slightly distad of Rs fork; m–cu always connecting base of MP with CuA. Crossveins between M and CuA and between CuA and CuP numerous. CuP straight, entering wing margin approximately at level of M fork. Hindwings with four longitudinal veins in anal area. Legs long and thin. First tarsomere long, almost twice as long as second; second tarsomere shorter than third. Cerci short, single-segmented." Line drawing from Liu et al. 2009, p. 189: References: N. D. Sinitshenkova. (1987). Istopicheskoe razvitie vesiyanok. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 221:1-142. Liu, Y.s., Sinitshenkova, N.D. & Ren, D. (2009). A revision of the Jurassic Stonefly Genera Dobbertiniopteryx Ansorge and Karanemoura Sinitshenkova (Insecta: Plecoptera), with the description of new species from the Daohugou locality, China. Paleontol. J. 43, 183–190. https://doi.org/10.1134/S0031030109020099
  7. oilshale

    Parakseneura sp

    From the album: Invertebrates

    Parakseneura sp Neuroptera Middle Jurassic Callovian - Oxfordian Daohugou Inner Mongolia PRC
  8. oilshale

    Prolyda elegantula Wang et al., 2016

    From the album: Invertebrates

    Prolyda elegantula Wang et al., 2016 Middle Jurassic Callovian to Oxfordian J iulongshan Formation Daohugou Inner Mongolia PRC
  9. From the album: Invertebrates

    Xyelotoma macroclada Gao, Ren & Shih, 2009 Middle Jurassic Callovian - Oxfordian Jiulongshan Formation Daohugou Inner Mongolia PR China
  10. oilshale

    Daohugocorixa vulcanica Zhang, 2010

    From the album: Invertebrates

    Daohugocorixa vulcanica Zhang, 2010 Jurassic Callovian - Oxfordian Jiulongshan Formation Daohugou Inner Mongolia China
  11. Taxonomy from Chen et al. 2019. Diagnosis from Wang et al., p. 1233: "Tegmen c. 3.5 times as long as wide, with apex widely rounded; basal cell c. 0.17 times as long as tegmen length, closed with anastomosis; common stalk ScR + M very short; branch ScRA c. 1.5 times as long as stem ScR; stem CuA at base distinctly convex, forking basad of claval apex; stigmal cell narrow, c. half as wide as radial cell." Line drawing from Wang et al. 2012, p. 1227: References: WANG, B., SZWEDO, J. & ZHANG, H. (2012). NEW JURASSIC CERCOPOIDEA FROM CHINA AND THEIR EVOLUTIONARY SIGNIFICANCE (INSECTA: HEMIPTERA) Palaeontology, Vol. 55, Part 6, 2012, pp. 1223–1243] Chen, J., Wang, B., Zheng, Y., Jiang, H., Jiang, T., Zhang, J. Q., An, B. Z. and Zhang, H. C. (2019). New fossil data and phylogenetic inferences shed light on the morphological disparity of Mesozoic Sinoalidae (Hemiptera, Cicadomorpha). Organisms Diversity & Evolution 19:287-302 [M. Clapham/M. Clapham]
  12. oilshale

    Parakseneura sp.

    Taxonomy from Fossilworks.com. Diagnosis for the genus from Yang et al. 2012, p. 4: 'Large neuropterans (forewing 50–75 mm long) with the following character states: labial palpi stout, relatively short; antennae stout, filiform, apparently much shorter than forewing length; two tibial spurs straight, shorter than basitarsus; claws big, strongly curved; in both wing, humeral veinlet well-developed, strongly recurrent, branched; presumed ScA short, fused with ScP within humeral area; membrane covered with dense, long hairs; RA (or ScP+RA) entering margin well before wing apex; subcostal crossveins numerous; radial crossveins irregularly spaced, not forming gradate series; in the forewing, MP, CuA, CuP dichotomously branched; presumed AA1+2 very short (found in Parakseneura gen. nov.); AA3+4, AP1+2, AP3+4 deeply forked; in hind wing, presumed AA1+2 very short (found in Pseudorapisma gen. nov.); proximal half of hind wings considerably wider than distal.' Determined by Dr. V. Makarkin, Academy of Sciences Vladivostok, Russia, as Paraksneura sp. It might be P. albadelta. Line drawing from Yang et al., 2012, p. 12: References: Yang Q, Makarkin VN, Winterton SL, Khramov AV, Ren D. (2012) A Remarkable New Family of Jurassic Insects (Neuroptera) with Primitive Wing Venation and Its Phylogenetic Position in Neuropterida. PLoS ONE 7(9): e44762. doi:10.1371/journal.pone.0044762.
  13. oilshale

    Platyxyela tenuis Zheng et al., 2021

    Taxonomy from Fossilworks.org. Etymology: The specific name tenuis, thin, feminine gender of the Latin, referring to the thinness and delicateness of the body. Diagnosis from Zheng et al. 2021, p. 150: "Forewing long (7.9 mm in length), ovipositor sheath as long as M+Cu and strongly narrowed toward acute apex." Line drawing from Zheng et al. 2021, p. 152: Identified by Prof. A. Rasnitsyn, Russian Academy of Sciences. References: Wang M., Shih C.K. & Ren D. 2012. Platyxyela gen. nov. (Hymenoptera, Xyelidae, Macroxyelinae) from the Middle Jurassic of China. Zootaxa 3456 (1): 82–588. Zheng Y., Hu H., Chen D., Chen J., Zhang H. & Rasnitsyn A.P. 2021. New fossil records of Xyelidae (Hymenoptera) from the Middle Jurassic of Inner Mongolia, China. European Journal of Taxonomy 733: 146–159.
  14. oilshale

    Parasinoala magnus Fu & Huang 2019

    Taxonomy from Fossilworks.com. Etymology: The specific epithet refers to the large size of the adult. Diagnosis from Fu & Huang (2019) p. 7:. Body length 15.3–16.1mm; tegmen with small dark bands on the median section, length/width ratio 3.3; ScP+RA relatively short, nearly 1.6 times longer than ScP+RA; RA with 2–3 branches; MP3+4 connecting CuA1 by cross-vein m-cu; hind wings, MP branching basal of CuA branching; cross-vein r-m basal of m-cu. Line drawings from Fu & Huang (2019) p. 8 and 9: Parasinoala magnus Scale bar 2 mm Left tegmen of Parasinoala magnus Right hind wings of Parasinoala minuta Scale bar 1 mm References: Yanzhe Fu & Diying Huang (2019): New sinoalids (Insecta: Hemiptera: Cercopoidea) from Middle to Upper Jurassic strata at Daohugou, Inner Mongolia, China, Alcheringa: An Australasian Journal of Palaeontology, DOI: 10.1080/03115518.2018.1528509
  15. Taxonomy from Fossilworks.org. Etymology: Derived from the Latin word formos- (beautiful) and Bittacus (a recent genus of Bittacidae). The species is named macularis after the presence of many maculae on wing. Diagnosis for the species †Formosibittacus macularis from Li et al. 2008, p. 42: "Sc very long, terminating at about three-fifth of the length of wing; sc-r at about two times its length before end of Sc; dark zones along cross-veins and darkened apex in wing membrane." Line drawing of body with wings, left forewing and left hindwing from Li et al., p. 41: References: Y. L. Li, D. Ren, and C. K. Shih. 2008. Two Middle Jurassic hanging-flies (Insecta: Mecoptera: Bittacidae) from northeast China. Zootaxa 1929: 38-46.
  16. oilshale

    Poljanka hirsuta Yang, Yao & Ren 2012

    Taxonomy from Fossilworks.org. Diagnosis from Yang et al. 2012, p. 38: "Scapus 1.50 times thicker than pedicelli, pedicelli two times thicker than flagellomeres, apical flagellomere swollen; femora about two times thicker than corresponding tibiae, hind tibiae 1.46–1.57 times longer than fore and mid tibiae, first tarsomere of fore legs 1.33 times longer than second, first tarsomere of mid legs 1.50 times longer than second, first tarsomere of hind legs 2.31 times longer than second; R1 reduced, vein M+CuA about 1.57 times longer than vein R, branches of M about 2.22–2.39 times longer than M." Line drawing from Yang et al. 2012, p. 39: References: G. Yang, Y. Yao & D. Ren (2012). A new species of Protopsyllidiidae (Hemiptera, Sternorrhyncha) from the Middle Jurassic of China. Zootaxa 3274: 36–42. G. Yang, Y. Yao & D. Ren (2013). Poljanka strigosa, a new species of Protopsyllidiidae (Hemiptera, Sternorrhyncha) from the Middle Jurassic of China. Alcheringa , 125–130. ISSN 0311-5518. Drohojowska, J., Szwedo, J., Żyła, D. et al. (2020). Fossils reshape the Sternorrhyncha evolutionary tree (Insecta, Hemiptera). Sci Rep 10, 11390. https://doi.org/10.1038/s41598-020-68220-x
  17. Chresmodidae are a family of insects that reigned from Middle Jurassic to Late Cretaceous. They are medium to large insects with very long legs and superficially resemble extant water striders. While they were common in water environments, like Daohugou, they are exceedingly rare in land environments like burmite. They are puzzling critters, since scientists suggest it is unlikely these weighty insects had the buoyancy necessary to be water surface predators like water striders (Gerridae, Hemiptera). These are samples of Daohugou Chresmodid fossils.
  18. Taxonomy from Mindat.org. Alternative name Netropanorpodes decorosus Sun, Ren & Shih 2007. Revised diagnosis from Cao et al. (2015), p. 9: "In forewings, Sc forks with three branches; R2+3 forks after the crossvein r1–r2+3; Cu1+M forks before the crossvein cu1–cu2. In hind wings, R2+3 forks before the crossvein r1–r2." Line drawing of the left forewing from Cao et al. 2015, p. 8: References: NAN LIU, YUNYUN ZHAO & REN (2010) Two new fossil species of Itaphlebia (Mecoptera: Nannochoristidae) from Jiulongshan Formation, Inner Mongolia, China. Zootaxa 2420: 37–45. CAO, Y., SHIH, C., BASHKUEV, A. & REN, D., XX.XX.XXXX. Revision and two new species of Itaphlebia (Nannochoristidae: Mecoptera) from the Middle Jurassic of Inner Mongolia, China. Alcheringa 40, XX–XX. ISSN 0311-5518. Cao Y, Lin X, Shih C, Ren D (2022) Two new species of Itaphlebia (Insecta, Mecoptera, Nannochoristidae) from the late Middle Jurassic of China. ZooKeys 1108: 175–188. https://doi.org/10.3897/zookeys.1108.85378
  19. Taxonomy from Fossilworks.org. Male scorpionfly. Species name reflects the characteristic pattern marking of the wings. Diagnosis from Soszyńska-Maj et al. 2019, p. 9: "Species differs from remaining congeneric by combination of the following characters: numerous oval-shaped small regular dark spots on the membrane all situated between veins spread evenly throughout the whole wings, more transparent areas than dark, Rs forks slightly beyond Mb forks in forewing, Rs1+2 almost twice as long and Rs3+4, basal part of M4b in forewing and M4 in hindwing strongly oblique and situated with cross-vein m-cu on one line." Line drawing from Soszyńska-Maj et al. 2019, p. 13 : References: Agnieszka Soszyńska-Maj, Wiesław Krzemiński, Katarzyna Kopeć, Yizi Cao, Ren, Wiesław Krzemiński & Katarzyna Kopeć (2019): New Middle Jurassic fossils shed light on the relationship of recent Panorpoidea (Insecta, Mecoptera), Historical Biology, DOI: 10.1080/08912963.2018.1564747
  20. Taxonomy from Fossilworks.org. Etymology. From “ellip- ”, which means elliptic and genus Ragio, referring to the elliptic wings. Gender: masculine. Diagnosis from Han et al. 2019, p.154: “Flagellum with 10 flagellemeres; proboscis long, labium fleshy, labella small. Wings elliptic and wide; R2+3 sinuate at the middle, and sharply up-curved distally; crossvein r-m intersecting the upper margin of d cell at basal one third (1/3); four medial veins present, bM3 and dM3 straight; anal cell closed before wing margin. Midtibiae with 1 spur.“ Line drawings from Han et al., 2019, p. 157 (scale bar equals 0.5mm): References: Han YE, Cai Y, Ren D, Wang Y. (2019). A new fossil snipe fly with long proboscis from the Middle Jurassic of Inner Mongolia, China (Diptera: Rhagionidae). Zootaxa. 30;4691(2):zootaxa.4691.2.4. Zhang, K., Yang, D. & Ren, D. (2008). A new genus and species of Middle Jurassic rhagionids from China (Diptera, Rhagionidae). Biologia, 63(1), 113-116. https://doi.org/10.2478/s11756-008-0012-4 Zhang, J. (2013). Snipe flies (Diptera: Rhagionidae) from the Daohugou Formation (Jurassic), Inner Mongolia, and the systematic position of related records in China. Palaeontology, 56, 1, 217–228. Zhang, K., Li, J., Yang, D., & Ren, D. (2009). A new species of Archirhagio Rohdendorf, 1938 from the Middle Jurassic of Inner Mongolia of China (Diptera: Archisargidae). Zootaxa, 1984, 61-65.
  21. oilshale

    Mesosciophila eucalla Zhang 2007

    Taxonomy from Fossilworks.org. Emended diagnosis for the genus from Zhang 2007, p. 298: "Medium-sized mesosciophilid gnats. Male body (including legs) covered with long, dense pubescence. Eyes large. Maxillary palps five-segmented, longer than head length. Antennae filiform, 16-segmented, with scapes and pedicels quadrate, flagellomeres cylindrical. Mesonotum convex. Scutellum clearly projecting. Venationally, Sc1 ending distad to level of Rs origin, Sc2 situated clearly basad to Rs origin; bRs longer than r-m; R1 slightly curved; both R1 and R4+5 divergent terminally; Rs furcated distad to fork of M1+2; R2+3 oblique; cell r moderately large, one-quarter to one-fifth of length of wing; stem of M not developed. Halteres light, with pubescence not visible. Femora, tibiae and first two tarsomeres with one or two rows of short setae." Line drawings from Zhang 2007, p. 300 (scale bars represent 1mm): A: Male Gnat C: Wing References: Junfeng Zhang (2007). New mesosciophilid gnats (Insecta: Diptera: Mesosciophilidae) in the Daohugou biota of Inner Mongolia, China, Cretaceous Research, Volume 28, Issue 2, Pages 297-301, ISSN 0195-6671, https://doi.org/10.1016/j.cretres.2006.05.007.
  22. oilshale

    Itaphlebia laeta Liu et al. 2010

    Taxonomy from Mindat.org. Diagnosis from Liu et al. 2010: "Costal area narrow. In forewing, Sc simple, terminating at C near pterostigma; M with 4 branches; Cu 1 and M forked before the cross-vein cu 1 -cu 2." Description from Liu et al. 2010: "Antenna filiform, incompletely preserved. Femur shorter than tibia. Tibial spurs not present. Setae irregularly arranged. Costal area narrow, one cross-vein c-sc between C and Sc. Sc simple, ending at C and extending to pterostigmal area. One short cross-vein sc 2 -r 1 before pterostigma. Rs arised at the same level to cross-vein csc. Rs with 4 branches. Length of R 2 two-thirds that of stem of R 2 + 3. Cross-vein r 1 -r 2 at level of pterostigma. Cross-vein r 3 -r 4 at level of cross-vein r 1 -r 2. Cross-vein r 5 -m 1 + 2 approximately at middle of wing length. Crossvein r 5 -m 1 proximal to cross-vein r 4 -r 5. One cross-vein r 4 -r 5 dividing cell R 4 into 2 cells. Conspicuous thyridium at fork of M, which has 4 branches. M 3 + 4 divided beyond the fork of M 1 + 2; cross-vein m 1 + 2 -m 3 between M 1 + 2 and M 3; cross-vein m-cu not straight, slightly S-shaped. Cu 1 coalesced with M for a short distance and separated from M before cross-vein cu 1 -cu 2. One cross-vein a 1 -a 2 connected 1 A and 2 A. 3 A absent. Female abdomen with 11 segments, segments 9–11 smaller than segment 8. One pair of cerci located at apex of abdomen. Right cercus with 3 segments. Third segment of left cercus not preserved. Basal segments fused with abdomen segment 11. Body length 7.8 mm, forewing length 8.2 mm, width 2.6 mm." Line drawing of the left forewing from Liu et al. 2010: Reference: NAN LIU, YUNYUN ZHAO & REN (2010) Two new fossil species of Itaphlebia (Mecoptera: Nannochoristidae) from Jiulongshan Formation, Inner Mongolia, China. Zootaxa 2420: 37–45.
  23. Taxonomy according to Hao et al 2009. Hao et al. 2009, p223: “Revised diagnosis: Body length: 6-13 mm: wing length 8.2-15.6 mm Sc ending at wing margin approximately the same as R4+5 forking into R4 and R5; R1 long, r-r at one -third of its length before the end of R1; R2+3 shorter than R3; R4+5 short, R4+5 fork six times longer than dR4+5. M1 smoothly curved, m-m joining close to M1+2 bifurcations. M3+4 with a little bend at m-m. CuA slightly sigmoid beyond m-cu; A1 strongly curved to wing margin in two-thirds of its length.” Wing line drawing from Hao et al. 2009, p.225: Identified by oilshale using Hao et al. 2009. References: Ren, D. & Krzemiński, W. 2002. Eoptychopteridae (Diptera) from the Middle Jurassic of China. Ann. Zoologica 52 (2): 207-210. Hao J., Ren, D. and Chungkun S. (2009) New Fossils of Eoptychopteridae (Diptera) from the Middle Jurassic of Northeast China Acta Geologica Sinica 83(2):222 – 228 DOI: 10.1111/j.1755-6724.2009.00022.x Liu, L., Shih, C. and Ren, D. (2012) Two new species of Ptychopteridae and Trichoceridae from the Middle Jurassic of northeastern China (Insecta: Diptera: Nematocera). Zootaxa 3501: 55-62, DOI: 10.5281/zenodo.282475
  24. oilshale

    Insect non det.

    From the album: Invertebrates

    Insect non det. (Mecoptera or Scorpionfly?) Middle Jurassic Daohugou Nei Mongol China
  25. Hi everyone, My friend bough some of Daohugou fossils for his museum and he asking to help for the ID. The only information that we got these fossils came from Nei Mongols, China. So do you have any idea about these fossil pls help us. Thanks for reading 1.
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