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Archaeohippus mannulus, sp. nov. Monroecreekian/Harrisonian terrestrial claystone Arikareean, late Oligocene/early Miocene Pinellas County, Florida On permanent display at the Florida Museum of Natural History, Gainesville, Fl. I discovered this particular specimen back in 95 while collecting fossils in a shallow creek. Initially thought to be a new species of Miohippus, it was sent to the Museum Of Natural History in Gainesville Fl. for further studies. In 2003 it was determined to be a new species of Archaeohippus rather than Miohippus.

Hunting between thunderstorms and deeper water. During the rest of the season, I note those places where I am finding fossils but have low water conditions...because of lower back issues, I generally refuse to hunt where I must bend over the screen. However, I do remember where such spots exist for days like today. An excellent day, some unusual, finds, a couple of megs, and then these: A odd bone, I have not previously found, but believe to be an Equus Splint bone: Another interesting fossil which I think most likely a large Sloth dermal scute. Finally, my best find of the day, a piece of jaw with a Hemiauchenia m3 molar in nice cream - brown colors.. These are really nice finds... but I was cherishing the end days of the 2017-2018 season with a friend on a day with sunshine in the morning and rain clouds later in the day. Does not get better than this... Jack

I've been wanting to get back to the Peace River since I first ventured out this fossil hunting season back in early February. Back then the water was over a foot higher and much colder--the air temps were in the mid-60s and the water was a chilly 62F. I decided this was a good day to test my new chest-high waders. I ventured into a spot I like to visit when I'm on this section of the Peace as it has some pretty coarse gravel. While it doesn't produce a lot of finds they tend to me more interesting. I waded out to the small patch of gravel at the leading edge of a sandbar but before I could reach the spot I found myself on tippy-toes trying to find a shallow path while the water rose to within an inch or so of the top of my waders. Somehow gathering more than my usual amount of common sense I decided to turn around rather than risk scuttling my new waders with a catastrophic flood. While searching around for another path to this gravel exposure I tried various approached though none were successful in attaining the desired location in the river that was tantalizingly close. While I walk the river I usually have my fiberglass probe (The Probulator 3000TM) in one hand pushing the tip into the sand with each step to test for any gravel crunch. Much to my surprise I was detecting a decent layer of gravel well downstream from the tiny outcrop on the leading upstream edge of the sandbar where I usually hunt. I have probed around this area before and only detected sand save for this one tiny area. Though I had found gravel in water that was a bit shallower I couldn't stay long as I had to be real careful to not bend over much while digging for gravel as it would have meant cold water down the waders. I couldn't lift as much with my legs and my lower back was soon very vocal in its complaint of the shifted workload. My upper body was also getting quite chilled as my long-sleeved shirt (good for solar protection) was getting soaked as usual but the brisk breeze was doing an efficient job at evaporative cooling quickly dropping my core body temp. I could only work for about 15-20 minute blocks before having to sit in the canoe and try to warm up my gradually numbing fingers. Instead, I conceded and made a mental not to return to investigate this increased exposure of gravel next time. I had hoped to get out last weekend but there was a bit of a cold front moving through Florida and the chance for rain shifted from late Saturday and on into Sunday to instead start mid-morning. I've been on the Peace when passing showers have opened up and spilled some precipitation down from above--not so bad on a warm day but not optimal for preserving core body temperature on a cooler day. Saint Patrick's Day weekend looked to have weather much more conducive to standing around half submerged in a river. The water temperature had risen to a relatively balmy 70F and the air temp was forecast to be an unseasonably warm 85F--unexpected as this was still technically winter with the Spring Equinox still two days hence. I had guests visiting and staying over on Friday night so it was not possible to get to the river on Saturday as I usually do but Sunday was clear. The morning started off a bit cool. I was up at 3:30am and out the door by 4:00am. The trip cross-state over the top of Lake Okeechobee and on into Arcadia was quiet (as it usually is that time of morning). I usually monitor the outside air temp on the car thermometer and watch it dip as I leave coastal Florida and cross over through its less populated center. I usually expect the temps to dip several degrees but this time I went from 67F as I left my neighborhood to the usual dip to near 60F. This time it continued even more and bottomed out at the nadir of 49F for a brief moment before rebounding into the 60s as we approached Arcadia. Most of the trip on two lane highway 70 was made more interesting by a thick coat of fog that approached white-out conditions a few times. It can be rather difficult to locate the road when the oncoming headlights of an approaching vehicle light swirling fog in an effect worthy of a Pink Floyd concert from the 1970's. We arrived without issue and went through the normal procedure of checking in at Canoe Outpost and riding the old blue school bus with canoe-laden trailer in tow to Brownville Park where we departed from the boat ramp into a white ethereal mist. For some reason the Earl Scruggs song "Foggy Mountain Breakdown" came to mind. https://www.youtube.com/watch?v=yQIJuu3N5EY Since we decided not to spend time at our normal spots further upstream, we soon left the rest of the canoes in our group as we headed off downstream into the dreamlike fog. The heavy mist also muffled sounds a bit so it was peacefully quiet and most befitting of its name. For some time we heard nothing more than the sounds of our paddles and a few species of birds calling. It was well worth the effort of the early departure just to experience this quiet time on the river. We saw some ducks who took to flight at our approach and enjoyed seeing some Great Blue Herons, Great Egrets, and Little Blue Herons hunting for a fishy breakfast along the banks of the river. There were lots of Cardinals, Gray Catbirds, and Belted Kingfishers in the trees that we would frequently spot flitting about or calling out to each other. Tammy mentioned that in all of the trips down the Peace that we had never seen an owl and she wished that for once she could see one here. Apparently, the officials at the Wish Granting Department had a light schedule this morning as, within 5 minutes of uttering this desire, she looked up into a tree at the edge of the river to spot a Barred Owl watching from its perch as we floated by. I pulled out the camera and we circled back for the photo. As we were leaving we saw the bird take flight. It is amazing how a bird this size can move on such stealthy wings as to be so utterly silent in flight. Our morning was made and I hadn't even broken out the shovel & sifting screen nor dipped foot into the water yet. I figured if this was a day for wishing that I'd put in my order for a reasonably complete mastodon tooth. These teeth are seemingly as fragile as mammoth teeth and mostly I've only found small but very distinctive (because of their thick pearlescent enamel in cross-section) chunks. I was fortunate enough to find a complete Colombian Mammoth tooth a few years back with John @Sacha but mastodon in anything but tiny fragments has so far eluded me. I made my wish and we continued to our destination. In time we made it down to my favorite sandbar and spent this entire trip focusing on seeing what this gravel had to offer. I couldn't determine if this was a new extended layer of fresh gravel that Hurricane Irma had chosen to spread out more evenly across the top of this sandbar or if the storm (and ensuing raging torrent) had stripped off a thick cap of sand uncovering an older previously-inaccessible gravel layer underneath. The water was lower that last time (and quite a bit warmer). No waders this time and after a few minutes for by legs to acclimate (read this as "becoming numb") I slowly worked my way into deeper water probing around with the Probulator and mapping out the extend of this newly expanded gravel. Tammy (being the wiser of the two) decided the morning was still too chilly for direct skin contact chose to sit in the canoe at the side of the river and drink from her thermos of hot tea. The river flow at this point in the river was nearly imperceptible (my tethered sifting screen occasionally floating slowly upstream rather than downstream). Being creative, Tammy decided that she could paddle out and position the canoe nearby and see what I was doing without the discomfort of standing in a river on a chilly morning before the sun was able to warm things up sufficiently. The sun finally burned off the morning fog and before long the sun's rays were counteracting the chilly water making the environmental conditions near optimal for standing around in a river. I got to work scouting out the extents of the gravel and picking some novel spots that I'd not dug before to see if I could detect some virgin gravel with worthy finds (nothing is worse than digging in spoil pile gravel with all of the work and none of the payoff). Before long some nice finds started appearing in the sifting screen. Because of the chunkiness of the gravel at this spot I choose to use my sifting screen with the 1/2" mesh rather than the finer 1/4" mesh screen. As a result, I found almost no smaller shark teeth (just a few larger ones that were not small enough to slip between the mesh back into the Peace). The gravel in this extended area was just as chunky as the former minor occurrence at the leading edge of the sandbar. It can bit a bit difficult to get a shovel into and a lot of wiggling around of the handle is necessary to slowly work the tip of the spade down between the stony chunks. Every now and then a shovel size chunk of matrix comes up on the shovel and threatens to sink the sifting screen with its bulk. I've learned to toss these behind me with reasonable care so as not to spray myself with the resultant depth-charge splash of chucking these bowling ball size chunks with too much vigor. There are some days on the Peace when even somewhat common items like horse teeth can be elusive. Today was not one of those days. The first horse tooth was a nice specimen of an upper Equus molar. It was soon followed by a nice lower Equus (the lowers are more thin and elongate to fit into the more narrow mandible). You can see the comparison of the two below.

Good evening, I've found this section a year ago and since then kept it in our garden. I took a second look at it tonight and thought it might be my first camel jaw piece. I found this along a creek in Travis county, thanks

Hi! I found a skull online, and think it looks great. But I'm not sure if it is a skull from a modern horse, or if it is a fossil skull. Can you see if it looks modern or very old? Cause I can't decide of my own if it is a "to good to be truth" skull.

Hey all, Any reason to believe that this horse leg might belong to anything besides Equus? It's from Ice Age Florida.

This complete metacarpal, a.k.a. cannon bone is an extremely rare find for North Carolina. Pleistocene mammals are uncommon and are mostly teeth. Being complete and undamaged it will be taking a prominent spot in my collection. The bone was found in a quarry containing mixed sediments of Pliocene Yorktown Formation, which is marine and a Pleistocene pebble lag. The odds are very very small that this would be from the Pliocene, so I am going on my gut feeling that this is from the Pleistocene. I also would like to send thanks to forum member @Fruitbat and @Harry Pristis for giving me a positive I.D. on this.

I believe this to be a horse metatarsal, most likely Equus. But am wondering since it was found in mixed Pleistocene / Pliocene sediments if it could be a Pliocene species. That is saying it is horse of course, you know Mr. Edd. It was found in a quarry in eastern NC that contains Pliocene Yorktown formation and a Pleistocene pebble lag. It was protruding from a sloped area that extended down into a shallow pit. The sediments pushed down were mixed Yorktown and the pebble lag. Mammal fossils are uncommon in North Carolina and they are usually teeth. Bones, especially complete are almost unheard of.

I found this tooth on the banks of the river Vilaine in Brittany in France. I believe it is from a horse. Can anyone identify it for me or let me know a possible age. Thanks, Jo

Hi, I recently found this bone on the shore of the Thames & wondered if anyone could help identifying it please. The bone is very dense & has a great colour & patination. From what I can gather it is a middle phalanx from a horse type creature?? What I'm trying to work out is possibly how old it could be & what type of horse it may have come from. Thanks here is the other side next to other objects I found

So I found a relatively small horse tooth. It is not hard to do. There are lots of Horse teeth in the Peace River, most of which are large but there are lots of small ones. A reason that this gets interesting is that Equus .sp is the most common horse species in the Peace River and goes back maybe 1.5 million years (there is some debate on the exact boundary line for the Pleistocene). Before Equus there were a great diversity of horse species, all smaller animals with smaller teeth. So whenever I find a small horse tooth: Is it Equus or is it an older smaller horse with smaller teeth? Here is a tooth from this week: So 26.5x11 Here are a couple of those smaller teeth from smaller horses: Here is a small Horse tooth found on the same day with Nannippus peninsulatus (above) that we agreed was still large enough to be Equus .sp All of the above teeth are lower jaw teeth (either m3 or p2 which look exactly the same) . So the question I have is: Is there a consensus on the minimal occlusal size (length and width) of Equus .sp lower jaw teeth? I have known examples of Equus (32x15) and Nannippus (19x9). Is a 26.5x11 tooth in the range of equus or not? All opinions and facts appreciated.

As some of you know , I volunteered to analyze a box of horse teeth for a fossil friend. Still working on it, but came up with a question for the Horse whisperers: Horses have a set of adult teeth that start around 4-5 inches and are worn down over years. New teeth are erupted at a maximum size. When the teeth wear out , the animal dies of starvation. I seem to have found a bunch of teeth that refute that understanding. I have 20-30 such non erupted teeth and many measure around 1.5 inches long and a few of them have roots. What am I missing .. How can the majority of unerupted teeth I have be so short? What am I missing ?

Horse Teeth 'Modern' horse teeth are very hypsodont (high-crowned) to deal with wear caused by eating gritty and/or fibrous foods like grasses. A mature horse may have as many as 44 teeth, which include: 12 incisors (6 upper and 6 lower) Canine teeth are usually absent in female horses but may be present in males. Cheek teeth (4 premolars and 3 molars per side) have very complex enamel patterns. The first premolars (upper and lower) in horses (sometimes called the 'wolf teeth') are vestigial and often absent. Upper cheek teeth (premolars and molars) can be recognized by the relatively square shape (except for the second premolar and third molar) when viewing the occlusal (chewing) surface. Lower cheek teeth (premolars and molars) can be recognized by the relatively rectangular shape when viewing the occlusal (chewing) surface (except for the second premolar and third molar). Horse 'foot' bones 'Modern' horses are monodactyl (one-toed). The metapodials (hand and foot bones) are reduced to a single unit on each leg. There are three 'toe bones' - phalanges (singular is phalanx) on each foot...phalanx I, phalanx II and phalanx III. The third phalanx is the 'hoof core' Unfortunately, I have never collected an intact phalanx III so I have not pictured one here. The astragalus (ankle bone) is only present on the hind legs.

I was poking around in a Texas site looking for my very own Mastadon Molar, with which CreekCrawler has so recently shamed us all. I did not find any elephant parts, but I did find a pretty cool Pliestocene Equus tooth - cheek tooth I think. Does anyone have a link to a good Pleistocene/Oligocene/Miocene Mammal tooth website or a good reference book? In Situ Reverse side Top pattern Huge Squali This is the most complete Globidens tooth I have found - about 75% there. Most of the back is missing unfortunately. One Crab that may have a pincher intact. I will explore the matrix later. Happy fossiling! Jon

These are a few of the pdf files (and a few Microsoft Word documents) that I've accumulated in my web browsing. MOST of these are hyperlinked to their source. If you want one that is not hyperlinked or if the link isn't working, e-mail me at joegallo1954@gmail.com and I'll be happy to send it to you. Please note that this list will be updated continuously as I find more available resources. All of these files are freely available on the Internet so there should be no copyright issues. Articles with author names in RED are new additions since May 26, 2018. Order Perissodactyla Family Equidae - The Horses Eocene Equidae Badiola, A., et al. (2002). First record of the genus Leptolophus Remy, 1965 (Mammalia, Perissodactyla) in the late Eocene (Priabonian) of Europe. Geodiversitas, 24(4). D'Ambrosia, A.R., et al. (2014). Stable isotope patterns found in early Eocene equid tooth rows of North America: Implications for reproductive behavior and paleoclimate. Palaeogeography, Palaeoclimatology, Palaeoecology, 414. Danilo, L., et al. (2013). A New Eocene Locality in Southern France Sheds Light on the Basal Radiation of Palaeotheriidae (Mammalia, Perissodactyla, Equoidea). Journal of Vertebrate Paleontology, 33(1). Franzen, J.L., C. Aurich and J. Habersetzer (2015). Description of a Well Preserved Fetus of the European Eocene Equoid Eurohippus messelensis. PLoS ONE, 10(10). Froehlich, D.J. (2002). Quo vadis eohippus? The systematics and taxonomy of the early Eocene equids (Perissodactyla). Zoological Journal of the Linnean Society, 134. Granger, W. (1908). A Revision of the American Eocene Horses. Bulletin American Museum of Natural History, Vol. XXIV, Article XV. Kitts, D.B. (1957). A Revision of the Genus Orohippus (Perissodactyla, Equidae). American Museum Novitates, Number 1864. Masciale, D.M. (2010). An Analysis of Anchitherine Equids Across the Eocene-Oligocene Boundary in the White River Group of the Western Great Plains. Masters Thesis - University of Nebraska. Rej, J.E. and S.G. Lucas (2016). Morphological Comparison of Two Early Eocene Horse Taxa: Minippus of New Mexico and Sifrhippus from Wyoming. In: Fossil Record 5. Sullivan, R.M. and S.G. Lucas (eds.), New Mexico Museum of Natural History and Science, Bulletin 74. Remy, J.A., G. Krasovec and B. Marandat (2016). A new species of Propalaeotherium (Palaeotheriidae, Perissodactyla, Mammalia) from the Middle Eocene locality of Aumelas (Herault, France). Palaeovertebrata, Vol.40(2). Rose, K.D., L.T. Holbrook and W.P. Luckett (2017). Deciduous premolars of Eocene Equidae and their phylogenetic significance. Historical Biology, 2017. Scott, W.B. (1891). On the Osteology of Mesohippus and Leptomeryx With Observations on the Modes and Factors of Evolution in the Mammalia. Journal of Morphology, Vol.5, Number 3. Secord, R., et al. (2012). Evolution of the Earliest Horses Driven by Climate Change in the Paleocene-Eocene Thermal Maximum. Science, Vol.335, Number 6071. Oligocene Equidae Farr, M.S. (1896). Notes on the Osteology of the White River Horses. The American Philosophical Society. Lambe, L.M. (1905). On The Tooth-Structure of Mesohippus westoni (Cope). The American Geologist, Vol.XXXV. McGrew, P.O. (1953). A New and Primitive Early Oligocene Horse from Trans-Pecos, Texas. Fieldiana Geology, Vol.10, Number 15. Osborn, H.F. (1904).New Oligocene Horses. Bulletin American Museum of Natural History, Volume XX, Article XIII. Prothero, D.R. and N. Shubin (1989). 10. The Evolution of Oligocene Horses. In: The Evolution of Perissodactyls. D.R. Prothero and R.M. Schoch (eds.), Oxford University Press. Scott, W.B. (1891). On the Osteology of Mesohippus and Leptomeryx With Observations on the Modes and Factors of Evolution in the Mammalia. Journal of Morphology, Vol.5, Number 3. Miocene Equidae Miocene Equidae - Africa/Middle East Bernor, R.L. (2007). The Latest Miocene Hipparionine (Equidae) from Lemudong'o, Kenya. Kirtlandia, Number 56. Bernor, R.L. (1985). Systematic and Evolutionary Relationships of the Hipparionine Horses from Maragheh, Iran (Late Miocene, Turolian Age). Palaeovertebrata, 15(4). Bernor, R.L. and R.S. Scott (2003). New interpretations of the systematics, biogeography and paleoecology of the Sahabi hipparions (latest Miocene)(Libya). Geodiversitas, 25(2). Bernor, R.L., N.T. Boaz and L. Rook (2012). Eurygnathohippus feibeli (Perissodactyla: Mammalia) from the Late Miocene of As Sahabi (Libya) and its Evolutionary and Biogeographic Significance. Bollettino della Societa Paleontologica Italiana, 51(1). Bernor, R.L., R.S. Scott and Y. Haile-Selassie (2005). A contribution to the evolutionary history of Ethiopian hipparionine horses (Mammalia, Equidae): morphometric evidence from the postcranial skeleton. Geodiversitas, 27(1). Bernor, R.L., T.M. Kaiser and S.V. Nelson (2004). The Oldest Ethiopian Hipparion (Equinae, Perissodactyla) from Chorora: Systematics, Paleodiet and Paleoclimate. Cour.Foursch.-Inst. Senckenberg, 246. Forsten, A. (1978). Hipparion and possible Iberian-North African Neogene connections. Ann.Zool. Fennici, 15. Franz-Odendaal, T.A., T.M. Kaiser and R.L. Bernor (2003). Systematics and dietary evaluation of a fossil equid from South Africa. South African Journal of Science, 99. Koufos, G.D. and T.D. Vlachou (2005). Equidae (Mammalia, Perissodactyla) from the late Miocene of Akkasdagi, Turkey. In: Geology, mammals and environments at Akkasdagi, late Miocene of Central Anatolia. Sen, S. (ed.), Geodiversitas, 27(4). Koufos, G.D. and D.S. Kostopoulos (1994). The late Miocene mammal localities of Kemiklitepe, Turkey: 3. Equidae. Bull.Mus.natl.Hist.nat., Paris, 4e ser.,16. Pickford, M. (2001). Equidae in the Ngorora Formation, Kenya, and the First Appearance of the Family in East Africa. Revista Española de Paleontología, 16(2). Scott, R.S. and M. Maga (2005). Paleoecology of the Akkaşdaği hipparions (Mammalia, Equidae), late Miocene of Turkey. Geodiversitas, 27(4). Scott, R.S., et al. (2003). 16. The Abundance of "Hipparion". In: The Geology and Paleontology of the Miocene Sinap Formation, Turkey. Fortelius, M., et al. (eds.), Columbia University Press. Watabe, M. and H. Nakaya (1991). Cranial Skeletons of Hipparion (Perissodactyla, Mammalia) from Maragheh (Turolian, Late Miocene), Northwest Iran. Memoirs of the Faculty of Science, Kyoto University, Series of Geol. & Mineral., Vol.LVI, Nunbers 1&2. Miocene Equidae - Asia/Malaysia/Pacific Islands Akhtar, M., et al. (2009). Hipparion from the Dhok Pathan Formation of the Middle Siwaliks, Pakistan. Punjab Univ.J.Zool., Vol.24(1-2). Babar, M.A., et al. (2016). Sivalhippus nagriensis (Equidae, Mammalia) from Dhok Pathan Formation of Siwaliks, Pakistan. The Journal of Animal & Plant Sciences, 26(4). Colbert, E.H. (1939). A New Anchitheriine Horse from the Tung Gur Formation of Mongolia. American Museum Novitates, Number 1019. Deng, T. (2006). Paleoecological comparison between late Miocene localities of China and Greece based on Hipparion faunas. Geodiversitas, 28(3). Deng, T. and X.-M. Wang (2004). Late Miocene Hipparion (Equidae, Mammalia) of Eastern Qaidam Basin in Qinghai, China. Vertebrata PalAsiatica, 42(4). Deng, T., et al. (2016). The Late Miocene Hipparion (Equidae, Perissodactyla) fossils from the Baogeda Ula, Inner Mongolia, China. Historical Biology, 28(1-2). Eronen, J.T., et al. (2014). Here be Dragons: Mesowear and tooth enamel isotopes of the classic Chinese "Hipparion" faunas from Baode, Shanxi Province, China. Ann.Zool. Fennici, 51. Forsten, A. (1982). The Taxonomic Status of the Miocene Horse Genus Sinohippus. Palaeontology, Vol. 25, Part 3. Hanif, M., et al. (2017). New Hipparionine (Perissodactyla) Fossils from Potwar Plateau of Pakistan. Pakistan J.Zool., Vol.49(5). Hou, S.-K., et al. (2007). New Materials of Sinohippus from Gansu and Nei Mongol, China. Vertebrata PalAsiatica, 45(3). Ikram, T., et al. (2016). Additional Fossils of Sivilhippus theobaldi (Mammalia, Equidae) from the Middle Siwaliks of Pakistan. Sci.Int.(Lahore), 28(3). Iqbal, M., et al (2009). Some New Remains of Hipparion from the Dhok Pathan Type Locality, Pakistan. The Journal of Animal & Plant Sciences, 19(3). Khan, M.A., et al. (2014). Some New Remains of Hipparionine (Equidae: Mammalia) from Dhok Pathan Upper Miocene, Northern Pakistan. Pakistan J.Zool., 46(2). Khan, M.A., et al. (2011). A New Collection of Hipparionine from the Type Locality of the Dhok Pathan Formation of the Middle Siwaliks. The Journal of Animal & Plant Sciences, 21(1). Li, Y., et al. (2017). Assessment of dental ontogeny in late Miocene hipparionines from the Lamagou fauna of Fugu, Shaanxi Province, China. PLoS ONE, 12(4). Miyata, K. and Y. Tomida (2010). Anchitherium (Mammalia, Perissodactyla, Equidae) from the Early Miocene Hiramaki Formation, Gifu Prefecture, Japan, And its Implication for the Early Diversification of Asian Anchitherium. J.Paleont., 84(4). Qiu, Z.-X. and J. Xie (1998). Notes on Parelasmotherium and Hipparion Fossils from Wangji, Dongxiang, Gansu. Vertebrata PalAsiatica, 36(1). Sun, B.-Y. (2013). The Miocene Hipparion (Equidae, Perissodactyla) from Shihuiba Locality, Lufeng, Yunnan. Vertebrata PalAsiatica, 51(2). Sun, B.-Y., et al. (2018). Sivalhippus ptychodus and Sivalhippus platyodus (Perissodactyla, Mammalia) from the Late Miocene of China. Rivista Italiana di Paleontologia e Stratigrafia, Vol.124(1). Tariq, M. and N. Jahan (2012). Paleoecology of Hipparion sp. (Equidae-Hipparionini) from latest Miocene of Padhri, northern Pakistan. Biologia (Pakistan), 58 (1&2). Watabe, M. (2011). Morphologic Comparison of Cranial and Postcranial Material of Chinese and Other Holarctic Hipparionine Horses. Vertebrata PalAsiatica, 49(3). Ye, J., W.Y. Wu and J. Meng (2005). Anchitherium from the Middle Miocene Halamagai Formation of Northern Junggar Basin, Xinjiang. Vertebrata PalAsiatica, 43(2). Miocene Equidae - Europe (including Siberia and Greenland) Bernor, R.L., H.-W. Mittmann and F. Rogl (1993). Systematics and Chronology of the Gotzendorf "Hipparion" (Late Miocene, Pannonian F, Vienna Basin). Ann.Naturhist.Mus. Wien, 95A. Bernor, R.L., et al. (2017). The Pannonian C hipparions from the Vienna Basin. Palaeogeography, Palaeoclimatology, Palaeoecology, 476. Bernor, R.L., et al. (2011). Systematics and Paleobiology of Hippotherium malpassii n.sp. (Equidae, Mammalia) from the latest Miocene of Baccinello V3 (Tuscany, Italy). Bollettino della Societa Paleontologica Italiana, 50(3). Bernor, R.L., et al. (2003). An Evaluation of the Late MN9 (Late Miocene, Vallesian Age), Hipparion Assemblage from Rudabanya (Hungary): Systematic Background, Functional Anatomy and Paleoecology. 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