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Fruitbat's Pdf Library - Class Maxillopoda: Barnacles, Fish Lice and Their Relatives

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These are a few of the pdf files (and a few Microsoft Word documents) that I've accumulated in my web browsing.

MOST of these are hyperlinked to their source.

If you want one that is not hyperlinked or if the link isn't working, e-mail me at joegallo1954@gmail.com and I'll be happy to send it to you.

Please note that this list will be updated continuously as I find more available resources. All of these files are freely available on the Internet so there should be no copyright issues.

Articles with author names in RED are new additions since July 17, 2017.


Phylum Arthropoda
Subphylum Crustacea


Class Maxillopoda - Barnacles and Their Relatives


Subclass Branchiura - Fish Lice and Cycloids


Dzik, J. (2008). Gill Structure and Relationships of the Triassic Cycloid Crustaceans. Journal of Morphology, 269.

Niko, S. and Y. Ibaraki (2011). First cyclid crustacean from East Asia. Jour.Geol.Soc. Japan, Vol.117, Number 4.

Schram, F.R., A.C. Boere and N. Thomas (2006). Cycloidea of the Mississippian Bear Gulch Limestone of Central Montana. Natural History Museum of Los Angeles County Contributions in Science Number 504. 

Schram, F.R., R. Vonk and C.H.J. Hof (1997). Mazon Creek Cycloidea. J.Paleont., 71(2). (202MB download)

Schweigert, G. (2007). Juracyclus posidoniae N.Gen. and Sp., the First Cycloid Arthropod from the Jurassic. J.Paleont., 81(1).

Schweigert, G. (2006). The first cycloid arthropod from the Late Jurassic. Zitteliana, A46.

van Bakel, B.W.M., et al. (2011). A new family, genus and species of cyclid (Crustacea, Branchiura, Cyclida) from mid-Cretaceous reefal deposits in northern Spain. Bulletin of the Mizunami Fossil Museum, Number 37.


Subclass Pentastomida - Tongue 'Worms'


Almeida, W.O., et al. (2008). Morphological support for the phylogenetic positioning of Pentastomida and related fossils. Biotemas, 21(3).

Lavrov, D.V., W.M. Brown and J.L. Boore (2004). Phylogenetic position of the Pentastomida and (pan)crustacean relationships. Proc.R.Soc.Lond. B, 271.

Siveter, D.J., et al. (2015). A 425-Million-Year-Old Silurian Pentastomid Parasitic on Copepods. Current Biology, 25.

Waloszek, D. and K.J. Muller (1994). Pentastomid parasites from the Lower Palaeozoic of Sweden. Transactions of the Royal Society of Edinburgh: Earth Sciences, 85.

Waloszek, D., J.E. Repetski and A. Maas (2006). A new Late Cambrian pentastomid and a review of the relationships of this parasitic group. Transactions of the Royal Society of Edinburgh: Earth Sciences, 96.

Waloszek, D., J.E. Repetski and K.J. Muller (1994). An exceptionally preserved parasitic arthropod, Heymonsicambria taylori n.sp. (Arthropoda incertae sedis: Pentastomida) from Cambrian-Ordovician boundary beds of Newfoundland, Canada. Canadian Journal of Earth Sciences, Vol.31, Number 11.


Subclass Thecostraca


Infraclass Cirripedia - Barnacles


Cirripedia - Africa/Middle East


Buckeridge, J.S., J.W.M. Jagt and R.P. Speijer (2008). Verruca punica, a new species of verrucomorph barnacle (Crustacea, Cirripedia, Thoracica) from the Lower Danian (Palaeocene) of Tunisia. Zootaxa, 1844.


Cirripedia - Antarctica


Zullo, V.A., R.M. Feldmann and L.A. Wiedman (1988). Balanomorph Cirripedia from the Eocene La Meseta Formation, Seymour Island, Antarctica. Geological Society of America, Memoir 169.

Cirripedia - Asia/Malaysia/Pacific Islands


Asami, K. and T. Yamaguchi (1997). Distribution of Living and Fossil Coral Barnacles (Cirripedia; Pyrogomatidae) in Japan. Sessile Organisms, 14(1).

Buckeridge, J.S. and J.A. Grant-Mackie (1985). A new scalpellid barnacle (Cirripedia: Thoracica) from the Lower Jurassic of New Caledonia. Geologie de la France, Number 1.

Nomura, S.-I. and H. Maeda (2008). Significance of autochthonous fossil barnacles from the Miocene Natori Group at the Moniwa-Goishi area, northeast Japan. Paleontological Research, Vol.12, Number 1.

Cirripedia - Australia/New Zealand


Beu, A.G. (1971). Further Fossil Whale Barnacles from New Zealand. New Zealand Journal of Geology and Geophysics, 14(4).

Buckeridge, J.S. (1991). Pachyscalpellum cramptoni: a new genus and species of lepadomorph cirripede from the Cretaceous of northern Hawke's Bay, New Zealand. Journal of the Royal Society of New Zealand, Vol.21, Number 1.

Buckeridge, J.S. (1984). Two new Tertiary scalpellid barnacles (Cirripedia: Thoracica) from the Chatham Islands, New Zealand. Journal of the Royal Society of New Zealand, Vol.14, Number 4.

Buckeridge, J.S. (1982). The barnacle subfamily Elminiinae - Two new subgenera and a new Miocene species from Victoria. Journal of the Royal Society of New Zealand, Vol.12, Number 4.

Buckeridge, J.S. (1979). The Fossil Barnacles (Cirripeda: Thoracica) of New Zealand and Australia (Part 1). Ph.D. Thesis - University of Aukland.

Buckeridge, J.S. (1979). The Fossil Barnacles (Cirripeda: Thoracica) of New Zealand and Australia (Part 2). Ph.D. Thesis - University of Aukland.

Buckeridge, J.S. (1975). The Significance of Cirripedes to the Paleoecology of Motutapu Island. Tane 21.

Fleming, C.A. (1959). A Pliocene Whale-Barnacle from Hawke's Bay, New Zealand. New Zealand Journal of Geology and Geophysics, 2(1).

Maxwell, P.A. (1968). A New Species of Lepas (Cirripedia, Lepadidae) from the Lower Miocene of New Zealand. New Zealand Journal of Geology and Geophysics, 11(1).


Cirripedia - Europe (including Greenland and Siberia)


Aguirre, J., et al. (2008). Densely packed concentrations of sessile barnacles (Cirripedia: Sessilia) from the Early Pliocene of SE Spain. Facies, 54.

Baluk, W. and A. Radwanski (1991). A new occurrence of fossil acrothoracican cirripedes: Trypetesa polonica sp.n. in hermitted gastropod shells from the Korytnica Basin (Middle Miocene; Holy Cross Mountains, Central Poland), and its bearing on behavioral evolution of the genus Trypetesa. Acta Geologica Polonica, Vol.41, Numbers 1-2.

Carriol, R.-P. and U. Menkveld-Gfeller (2010). Balanoidea (Crustacea, Cirripedia) from the Upper Marine Molasse (Early Miocene) of the Bern area, Switzerland. Contributions to Natural History, Number 13.

Carriol, R.-P. and S. Schneider (2008). A new Concavinae (Cirripedia, Chesaconcavus) from the Late Burdigalian (Miocene) of Lower Bavaria (Germany). N.Jb.Geol.Palaont. Abh., 248/3.

Carriol, R.-P. and I. Dieni (2005). Three new Verrucidae (Crustacea: Cirripedia) from the Upper Eocene of the Euganean Hills (Northeast Italy). Geobios, 38.

Collareta, A., et al. (2016). A new whale barnacle from the early Pleistocene of Italy suggest an ancient right whale breeding ground in the Mediterranean. C.R. Palevol, xxx. (Article in Press)

Collins, J.S.H. and A. Radwanski (1982). Scalpellid cirripedes from the Upper Cretaceous chalk of Mielnik (eastern Poland). Acta Geologica Polonica, Vol.32, Numbers 1-2.

Darwin, C. (1858). A Monograph on the Fossil Balanidae and Verucidae of Great Britain. The Palaeontographical Society. (185 pages)

Darwin, C. (1851). A Monograph on the Fossil Lepadidae, or, Pedunculated Cirripedes of Great Britain. The Palaeontographical Society. (186 pages)

Dermitzakis, M.D., et al. (2009). Lower Pliocene (Zanclean) regressive sequence of Rafina near Pikermi in Attica, Greece: a spectacular locality of mass-aggregated giant balanid cirripedes. Hellenic Journal of Geosciences, Vol.44.

Dominici, S., et al. (2011). Large kings with small crowns: a Mediterranean Pleistocene whale barnacle. Bollettino della Societa Paleontologica Italiana, 50(2).

Doyle, P., et al. (1997). Miocene barnacle assemblages from southern Spain and their palaeoenvironmental significance. Lethaia, Vol. 29.

Gale, A.S. (2014). New cirripedes (Crustacea, Thoracica) from the Jurassic and Cretaceous of the United Kingdom. Proceedings of the Geologists' Association, 125.

Gale, A.S. and G. Schweigert (2015). A New Phosphatic-Shelled Cirripede (Crustacea, Thoracica) from the Lower Jurassic (Toarcian) of Germany - The Oldest Epiplanktonic Barnacle. Palaeontology.

Gale, A.S. and A.M. Sorensen (2015). Taxonomy and palaeoecology of thoracican cirripedes (Crustacea) from a Campanian rocky shoreline at Ivo Klack, southern Sweden. Cretaceous Research, 54.

Gale, A.S. and A.M. Sorensen (2014). Origin of the balanomorph barnacles (Crustacea, Cirripedia, Thoracica): New evidence from the Late Cretaceous (Campanian) of Sweden. Journal of Systematic Palaeontology.

Hodges, P. (1995). A New Barnacle from the Lower Jurassic of South Wales. Palaeontology, Vol.37, Part 4.

Jagt, J.W.M., T.D. Zonova and E.A. Jagt-Yazykova (2007). A review of the brachylepadomorph cirripede genus Pycnolepas, including the first record of an Early Cretaceous species from the Russian Far East. Zootaxa, 1545.

Jagt, J.W.M., et al. (2008). A new record of the Late Cretaceous cirripede Eoverruca hewitti (Verrucomorpha, Proverrucidae) from southern Poland. Zootaxa, 1671.

Koci, T. and M.K. Veselska (2013). The First Recorded Occurrence of Smilium ? parvulum Withers, 1914 (Cirripedia, Thoracica) from the Bohemian Cretaceous Basin (The Czech Republic). Acta Musei Nationalis Pragae, Vol.69, Numbers 3-4.

Koci, T., et al. (2015). Late Jurassic-Early Cretaceous stalked barnacles (Cirripedia, genus Eolepas Withers, 1928) from Stramberk, Moravia (Czech Republic). N.Jb.Geol.Paleontol. Abh., 275/2.

Martin-Gonzalez, E., et al. (2012). First record of a tropical shallow water barnacle Tetraclita sp. (Cirripedia: Tetraclitoidea) from the middle Neogene of the Canary Islands. Vieraea, Vol.40.

Nagler, C., et al. (2017). Litholepas klausreschi gen. et sp.nov., a new neolepadine barnacle (Cirripedia, Thoracica) on a sponge from the Upper Jurassic lithographic limestones of southern Germany. N.Jb.Geol.Palaont. Abh., 284/1.

Radwanska, U. and A. Radwanski (2008). Eco-taphonomy of mass-aggregated giant balanids Concavus (Concavus) concavus (Darwin, 1854) from the Lower Pliocene (Zanclean) of Rafina near Pikermi (Attica, Greece). Acta Geologica Polonica, Vol.58, Number 1.

Veseleska, M.K. and T. Koci (2015). Taphonomy, distribution and palaeoecology of stalked barnacles (Crustacea, Cirripedia) from the Bohemian Cretaceous Basin, Czech Republic. Geology, Geophysics & Envronment, Vol.41(1).

Winkelmann, K., et al. (2010). Zullobalanus santamariaensis sp.nov., a new late Miocene barnacle species of the family Archaeobalanidae (Cirripedia: Thoracica), from the Azores. Zootaxa, 2680.

Cirripedia - North America


Buckeridge, J.S. and K.L. Finger (2001). First Record of a Fossil Verrucid Barnacle in California - Verruca digitali sp.nov. (Cirripedia: Thoracica) from the Late Miocene. Journal of Crustacean Biology, 21(2).

Ross, A. and W.A. Newman (1967). Eocene Balanidae of Florida, Including a New Genus and Species with a Unique Plan of "Turtle-Barnacle" Organization. American Museum Novitates, Number 2288.

Whitfield, R.P. (1889). Article V. Description of a new form of fossil, Balanoid Cirripede, from the Marcellus Shale of New York. Bulletin American Museum of Natural History, Vol.II, Number 2.

Zullo, V.A. (1979). Thoracican Cirripedia of the Lower Pliocene Pancho Rico Formation, Salinas Valley, Monterey County, California. Contributitons in Science - Natural History Museum of Los Angeles County, Number 303.


Cirripedia - South America/Central America/Caribbean


Bianucci, G., et al. (2006). Palaeoecology and taphonomy of an extraordinary whale barnacle accumulation from the Plio-Pleistocene of Ecuador. Palaeogeography, Palaeoclimatology, Palaeoecology, 242.

Carriol, R.-P. and S. Schneider (2013). New species of fossil sessile barnacles (Verrucidae, Tetraclitidae) from the Pliocene of central Chile. N.Jb.Geol.Palaont. Abh., 270/3.

Ifrim, C., F.J. Vega and W. Stinnesbeck (2011). Epizoic Stramented Cirripedes on Ammonites from Late Cretaceous Platy Limestones in Mexico. Journal of Paleontology, 85(3).


General Cirripedia

Collareta, A., M. Bosselaers and G. Bianucci (2016). Jumping from Turtles to Whales: A Pliocene Fossil Record Depicts an Ancient Dispersal of Chelonibia on Mysticetes. Revista Italiana di Paleontologia y Stratigrafia, Vol.22(2).

Gale, A.S. (2014). Origin and phylogeny of verrucomorph barnacles (Crustacea, Cirripedia, Thoracica). Journal of Systematic Palaeontology, 13(9).

Harzhauser, M., W.A. Newman and P. Grunert (2011). A new Early Miocene barnacle lineage and the roots of sea-turtle fouling Chelonibiidae (Cirripedia, Balanomorpha). Journal of Systematic Palaeontology, Vol.9, Issue 4.

Hayashi, R., et al. (2013). Phylogenetic position and evolutionary history of the turtle and whale barnacles (Cirripedia: Balanomorpha: Coronuloidea). Molecular Phylogenetics and Evolution, 67.

Jagt, J.W.M. and J.S. Buckeridge (2005). A case of homonymy in fossil verrucid barnacles: Verruca withersi (Crustacea, Thoracica). Scripta Geologica, 130.

Klompmaker, A.A., et al. (2015). The fossil record of drilling predation on barnacles. Palaeogeography, Palaeoclimatology, Palaeoecology, 426.

Perez-Losada, M., J.T. Hoeg and K.A. Crandall (2004). Unraveling the Evolutionary Radiation of the Thoracican Barnacles Using Molecular and Morphological Evidence: A Comparison of Several Divergence Time Estimation Approaches. Syst.Biol., 53(2).
Perez-Losada, M., et al. (2008). The tempo and mode of barnacle evolution. Molecular Phylogenetics and Evolution, 46.

Seilacher, A. (1969). Paleoecology of Boring Barnacles. Am.Zool., 9.


General Maxillopoda


Newman, W.A. (1992). Origin of Maxillopoda. Acta Zoologica (Stockholm), Vol.73, Number 5.


Copepodoida (Unranked)


Haug, J.T., et al. (2011). External Morphology and Post-Embryonic Development of Derocheilocaris remanei (Mystacocarida) Revisited, With a Comparison to the Cambrian Taxon Skara. Journal of Crustacean Biology, 31(4).

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Created and updated June 9, 2015.


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Updated August 26, 2015.


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Updated August 18, 2016.


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Fixed! Thanks doushantuo!


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Updated February 27, 2017.



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A word of caution: The Schram /Vonk piece is 202 Mb

Good find,Joe.


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Duly noted, doushantuo...thanks!


Updated July 17, 2017.



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