Jump to content
Sign in to follow this  

Fruitbat's Pdf Library - Subdivision Batoidea (Rays, Skates and Sawfishes)

Recommended Posts


These are a few of the pdf files (and a few Microsoft Word documents) that I've accumulated in my web browsing.

MOST of these are hyperlinked to their source.

If you want one that is not hyperlinked or if the link isn't working, e-mail me at joegallo1954@gmail.com and I'll be happy to send it to you.


Please note that this list will be updated continuously as I find more available resources. All of these files are freely available on the Internet so there should be no copyright issues.


Articles with author names in RED are new additions since May 25, 2018.



Class Chondrichthyes - The Cartilaginous Fishes

Subclass Elasmobranchii


Infraclass Euselachii (Sharks and Rays)


Division Neoselachii


Superorder Batoidea - Rays, Skates and Sawfishes


Order Myliobatiformes - Rays


Myliobatiformes - Africa/Middle East


Cappetta, H. (1984). Discovery of the Genus Gymnura (Batomorphii, Myliobatiformes) in the Thanetian of the Ouled Abdoun, Morocco. Observations on the Dentition of some Modern Species. Geobios, 17. (Plates not included)

Claeson, K.M., C.J. Underwood and D.J. Ward (2013). Ɨ Tingitanius tenuimandibulus, a New Platyrhinid Batoid from the Turonian (Cretaceous) of Morocco and the Cretaceous Radiation of the Platyrhinidae. Journal of Vertebrate Paleontology, 33(5).

Claeson, K.M., et al. (2010). First Mesozoic record of the stingray Myliobatis wurnoensis from the late Cretaceous of Mali and a phylogenetic study of Myliobatidae (Batoidea) incorporating dental characters. Acta Palaeontologica Polonica, 55(4).


Myliobatiformes - Asia/Malaysia/Pacific Islands


Adnet, S., et al. (2008). First myliobatiform teeth (Elasmobranchii, Neoselachii) from the Pondaung Formation (late middle Eocene) of Central Myanmar. N.Jb.Geol.Palaont. Abh., Vol.247/3.

Hatai, K. M. Murata and K. Masuda (1965). 485. Sting Ray and Eagle Ray from the Tatsunokuchi Formation (Pliocene) in Sendai City, Miyagi Prefecture, Japan. Trans.Proc.Palaeont.Soc. Japan, N.S., Number 57.

Marrama, G., et al. (2018). Anatomy, relationships and palaeobiogeographic implications of the first Neogene holomorphic stingray (Myliobatiformes: Dasyatidae) from the early Miocene of Sulawesi, Indonesia, SE Asia. Zoological Journal of the Linnean Society, xx.

Mishra, V.P. (1980). A New Species of Myliobatis and Some Shark Teeth from the Middle Eocene of Kutch, Western India. Journal of the Palaeontological Society of India, Vols. 23 & 24.


Myliobatiformes - Australia/New Zealand


Myliobatiformes - Europe (including Greenland and Siberia)


Antunes, M.T. and A.C. Balbino (2006). Latest Miocene Myliobatids (Batoidea, Selachii) from the Alvalade Basin, Portugal. Cainozoic Research, 4(1-2).

Bor, T.J. (1990). A New Species of Mobulid Ray (Elasmobranchii, Mobulidae) from the Oligocene of Belgium. Contr. Tert. Quatern. Geol., 27(2-3).

Hovestadt, D. and M. Hovestadt-Euler (2010). Urobatis molleni Nov.Sp. (Chondrichthyes, Myliobatiformes, Urolophidae) in the Eocene of Belgium. Geominpal Belgica, 1(3).


Myliobatiformes - North America


Cicimurri, D.J. (2010). On the dentition of Meridiana convexa Case (Myliobatoidea), an extinct Early Eocene ray from the United States. Cainozoic Research, 7(1-2).

Cicimurri, D.J. and J.A. Ebersole (2015). Two new species of Pseudaetobatus Capetta, 1986 (Batoidei, Myliobatidae) from the southeastern United States. Palaeontologia Electronica, 18.1.15A.

Cook, T.D., et al. (2014). A New Genus and Species of Freshwater Stingray (Myliobatiformes, Dasyatoidea) from the Latest Middle Eocene of Utah, U.S.A. Journal of Paleontology, 88(3).

De Carvalho, M.R., J.G. Maisey and L. Grande (2004). Freshwater Stingrays of the Green River Formation of Wyoming (Early Eocene), With the Description of a New Genus and Species and an Analysis of Its Phylogenetic Relationships (Chondrichthyes: Myliobatiformes). Bulletin of the American Museum of Natural History, Number 284.

de Santana, F.R., D.J. Cicimurri and J.A. Barbosa (2011). New Material of Apocopodon sericeus Cope, 1886 (Myliobatiformes, Myliobatidae) from the Paraiba Basin (Northeastern Brazil) and South Carolina (USA) With a Reanalysis of the Species. PalArch's Journal of Vertebrate Palaeontology, 8(6).

McNulty, C.L. (1964). Hypolophid Teeth from the Woodbine Formation, Tarrant County, Texas. Eclogae Geologicae Helvetiae, 57(2).


Myliobatiformes - South America/Central America/Caribbean


Adnet, S., R.S. Gismondi and P.-O. Antoine (2013). Comparisons of dental morphology in river stingrays (Chondrichthyes: Potamotrygonidae) with new fossils from the middle Eocene of Peruvian Amazonia rekindle debate on their evolution. Naturwissenschaften.

Cione, A.L., M. Tejedor and F.J. Goin (2012). A new species of the rare batomorph genus Hypolophodon (? latest Cretaceous to earliest Paleocene, Argentina). N.Jb.Geol.Palaont. Abh., 267/1.

de Santana, F.R., D.J. Cicimurri and J.A. Barbosa (2011). New Material of Apocopodon sericeus Cope, 1886 (Myliobatiformes, Myliobatidae) from the Paraiba Basin (Northeastern Brazil) and South Carolina (USA) With a Reanalysis of the Species. PalArch's Journal of Vertebrate Palaeontology, 8(6).


General Myliobatiformes


Adnet, S., et al. (2018). Teeth, fossil record and evolutionary history of the cowtail stingray Pastinachus Ruppell, 1829. Historical Biology, 2018.

Adnet, S., et al. (2012). Evolutionary history of the devilrays (Chondrichthyes: Myliobatiformes) from fossil and morphological inference. Zoological Journal of the Linnean Society, 166.

Bertozzi, T., M.S.Y. Lee and S.C. Donnellan (2016). Stingray diversification across the end-Cretaceous extinctions. Memoirs of Museum Victoria, 74.


Order Rajiformes - Skates


De Carvalho, M.R. (2004). A Late Cretaceous thornback ray from southern Italy, with a phylogenetic reappraisal of the Platyrhinidae (Chondrichthys: Batoidea).In: Mesozoic Fishes 3 - Systematics, Paleoenvironments and Biodiversity. Arratia, G. and A. Tintori, eds.

Herman, J., et al. (1994). Part B: Batomorphii No.1A: Order Rajiformes - Suborder Rajoidei- Family: Rajidae. In: Contributions to the comparative morphology of teeth and other ichthyodorulites in living supra-specific taxa of Chondrichthyan fishes. Stehmann, M. (ed.), Bulletin De L'Institut Royal Des Sciences Naturelles de Belgique, Biologie 64. (Note: While this article deals with living types of rays, it contains excellent references to ray tooth morphology and has a large number of pictures of ray teeth for comparative purposes!  Thanks to doushantuo for pointing this one out!)

Long, D.J. (1994). Quaternary colonization of Paleogene persistence?: historical biogeography of skates (Chondrichthyes: Rajidae) in the Antarctic ichthyofauna. Paleobiology, 20(2).

Siverson, M. and H. Cappetta (2001). A Skate in the Lowermost Maastrichtian of Southern Sweden. Palaeontology, Vol.44, Part 3.

Vullo, R. and D. Néraudeau (2008). When the "primitive" shark Tribodus (Hybodontiformes) meets the "modern" ray Pseudohypolophus (Rajiformes): the unique co-occurrence of these two durophagous Cretaceous selachians in Charentes (SW France). Acta Geologica Polonica, Vol.58, Number 2.


Order Rhinobatiformes - Guitarfishes


Bor, T.J. (1983). A New Species of Rhinobatos (Elasmobranchii, Batomorphii) from the Upper Maastrichtian of the Netherlands and Belgium. Geologie en Mijnbouw.

Brito, P.M., M.E.C. Leal and V. Gallo (2013). A New Lower Cretaceous Guitarfish (Chondrichthyes, Batoidea) from the Santana Formation, Northeastern Brazil. Boletim do Museo Nacional, Number 76.

Claeson, K.M., D.J. Ward and C.J. Underwood (2010). 3-D digital imaging of a concretion-preserved batoid (Chondrichthyes, Elasmobranchii) from the Turonian (Upper Cretaceous) of Morocco. C.R. Palevol, 9.

Everhart, M.J. (2007). New stratigraphic records (Albian-Campanian) of Rhinobatos sp. (Chondrichthyes; Rajiformes) from the Cretaceous of Kansas. Transactions of the Kansas Academy of Science, Vol.110, Numbers 3/4.

Langston, W. (1970). A Fossil Ray, Possibly Myledaphus (Elasmobranchii: Batoidea) from the Late Cretaceous Oldman Formation of Western Canada. National Museums of Canada, Publications in Palaeontology, Number 6.

Maisey, J.G. (1976). The Jurassic Selachian Fish Protospinax Woodward. Palaeontology, Vol.19, Part 4.


Order Torpediniformes - Electric Rays


Marrama, G., et al. (2018). Revision of Eocene electric rays (Torpediniformes, Batomorphii) from the Bolca Konservat-Lagerstätte, Italy, reveals the first fossil embryo in situ in marine batoids and provides new insights into the origin of trophic novelties in coral reef fishes. Journal of Systematic Palaeontology, Vol.16, Number 14.  (Thanks to doushantuo for finding this one!)

Ward, D.J. (1983). Additions to the fish fauna of the English Palaeogene. 4. A new batoid genus from the Bracklesham Group of Selsea, Sussex. Tertiary Research, 5(2).


General Batoidea


Ashliman, N.C., et al. (2012). Body plan convergence in the evolution of skates and rays (Chondrichthyes: Batoidea). Molecular Phylogenetics and Evolution, 63.

Dean, M.N., J.J. Bizarro and A.P. Summers (2007). The evolution of cranial design, diet and feeding mechanisms in batoid fishes. Integrative and Comparative Biology, Vol.47, Number 1.

Deynat, P.P. and P. Brito (1994). Revision of the Dermal Tubercles of Rays (Chondrichthyes: Batoidea) from the Parana Basin, Tertiary of South America. Annales de Paleontologie (Vert.-Invert.), 80(4).

Guinot, G., et al. (2012). Batoids (Elasmobranchii: Batomorphii) from the British and French Late Cretaceous. Journal of Systematic Palaeontology, Vol.10, Issue 3.

Hovestadt, D.C . and M. Hovestadt-Euler (1995). Additions to the fauna of the Boom Clay Formation of Belgium (Rupelian, Oligocene). Taxonomic adjustments on the Scyliorhinidae and Rajoidei, discovery of a dasyatid species (Pisces, Chondrichthyes) and of a cucrulionid species (Insecta, Coleoptera). Belgian Geological Survey, Professional Paper 278.

Kachacha, G., et al. (2017). Revision of the fossil batomorphs from the Cretaceous of Lebanon, and their impact on our understanding of the early step of the evolution of the clade. Research & Knowledge, Vol.3, Number 2.

Lowemark, L. (2015). Evidence for targeted elasmobranch predation on thalassinidean shrimp in the Miocene Taliao Formation, NE Taiwan. Lethaia, Vol.48.

Sharma, K.M. and R. Patnaik (2013). Additional Fossil Batoids (Skates and Rays) from the Miocene Deposits of Baripada Beds, Mayurbhang District, Orissa, India. Earth Science India, Vol.6 (IV).

Tiwari, R.P. and V.Z. Ralte (2012). Fossil batoid and teleost fish remains from Bhuban Formation (Lower to Middle Miocene), Surma Group, Aizawl, Mizoram. Current Science, Vol.103, Number 6.

Underwood, C.J., M.A. Kolmann and D.J. Ward (2017). Paleogene Origin of Planktivory in the Batoidea. Journal of Vertebrate Paleontology, e1293068.

Underwood, C.J., et al. (2015). Development and Evolution of Dentition Pattern and Tooth Order in the Skates and Rays (Batoidea: Chondrichthyes). PLoS ONE, 10(4).

van Netten, H.H. and J.W.F. Reumer (2009). Bite marks on early Holocene Tursiops truncatus fossils from the North Sea indicate scavenging by rays (Chondrichthyes, Rajidae). Netherlands Journal of Geosciences, 88-3.

Wing, E. (1966). Fossil Skates and Rays of Florida. The Plaster Jacket, Number 2. (Thanks to Nimravus for pointing me to this one!)


Clade Pristiorajea - Rhinopristiformes (Sawfishes) and Sclerorhynchiformes


Order Rhinopristiformes - Sawfishes


Carrillo-Briceno, J.D., et al. (2015). Sawfishes and Other Elasmobranch Assemblages from the Mio-Pliocene of the South Caribbean (Urumaco Sequence, Northwestern Venezuela). PLoS ONE, 10(10).

Collareta, A., S. Casati and A. Di Cencio (2017). A Pristid Sawfish from the Lower Pliocene of Lucciolabella (Radicofani Basin, Tuscany, Central Italy). Atti Soc.Tosc.Sci.Nat.Mus. Serie A, 124.

Deynat, P.P. (2005). New data on the systematics and interrelationships of sawfishes (Elasmobranchii, Batoidea, Pristiformes). Journal of Fish Biology, 66.

Farres, F. and H.L. Fierstine (2009). First record of the extinct sawfish Propristis schweinfurthi Dames, 1883 (Batoidea: Pristiformes: Pristidae) from the middle Eocene of Spain. Palaontologische Zeitschrift.

Schaeffer, B. (1963). Cretaceous Fishes from Bolivia, with Comments on Pristid Evolution. American Museum Novitates, Number 2159.


Order Sclerorhynchiformes


Arambourg, C. (1940). The Group of the Ganopristines. Bulletin de la Societe Geologique de France, Ser.5, 10. (Plates not included.)

Delgadillo-Escobar, A.A., et al. (2015). The first record of Onchosaurus (†Sclerorhynchidae) from the Late Cretaceous of northern Mexico. Boletín de la Sociedad Geológica Mexicana, Vol.67, Number 1.

Keyes, I.W. (1977). Records of the Northern Hemisphere Cretaceous Sawfish Genus Onchopristis (Order Batoidea) from New Zealand. New Zealand Journal of Geology & Geophysics, Vol.20, Number 2.

Kirkland, J.I. and M.C. Aguillon-Martinez (2002). Schizorhiza: a unique sawfish paradigm from the Difunta Group, Coahuila, Mexico. Revista Mexicana de Ciencias Geologicas, Vol.19, Number 1.

Knight, J.L., D.J. Cicimurri, and R.W. Purdy (2007). New Western Hemisphere Occurrences of Schizorhiza Weiler, 1930 and Eotorpedo White, 1934 (Chondrichthyes, Batomorphii). Paludicola, 6(3).

Kriwet, J. and S. Klug (2012). Presence of the extinct sawfish Onchosaurus (Neoselachii, Sclerorhynchiformes) in the Late Cretaceous of Peru with a review of the genus. Journal of South American Earth Sciences, 39.

Kriwet, J. and K. Kussius (2001). Paleobiology and Paleobiogeography of Sclerorhynchid Sawfishes (Chondrichthyes, Batomorphi). Revista Española de Paleontología, no. extraordinario.

Pereira, A.A. and M.A. Medeiros (2008). A New Sclerorhynchiform (Elasmobranchii) from the Middle Cretaceous of Brazil. Rev.bras.paleontol., 11(3).

Smith, M.M., et al. (2015). Early development of rostrum saw-teeth in a fossil ray tests classical theories of the evolution of vertebrate dentitions. Proc.R.Soc. B, 282.

Suárez, M.E. and H. Cappetta (2004). Sclerorhynchid teeth (Neoselachii, Sclerorhynchidae) from the Late Cretaceous of the Quiriquina Formation, central Chile. Andean Geology, Vol.31, Number 1.

Underwood, C., M.M. Smith and Z. Johanson (2015). Sclerorhynchus atavus and the convergent evolution of rostrum-bearing chondrichthyans. In: Arthur Smith Woodward: His Life and Influence of Modern Vertebrate Palaeontology. Johanson, Z., et al. (eds.), Geological Society, London, Special Publications, 430.


General Pristiorajea


Welten, M., et al. (2015). Evolutionary origins and development of saw-teeth on the sawfish and sawshark rostrum (Elasmobranchii; Chondrichthyes). R.Soc.Open Sci., 12.

Wueringer, B.E., L. Squire and S.P. Collin (2009). The biology of extinct and extant sawfish (Batoidea: Sclerorhynchidae and Pristidae). Rev. Fish Biol. Fisheries, 19.


Superorder Selachimorpha 


Order Pristiophoriformes - Sawsharks (not Sawfish)


Applegate, S. and T. Uyeno (1968). The First Discovery of a Fossil Tooth belonging to the Shark Genus Heptranchias, with a New Pristiophorus Spine, Both from the Oligocene of Japan. Bull.Nat.Sci.Mus. Tokyo, 11(2).

Keyes, I.W. (1982). The Cenozoic sawshark Pristiophorus lanceolatus (Davis) (Order Selachii) of New Zealand and Australia, with a review of the phylogeny and distribution of world fossil and extant pristiophoridae. New Zealand Journal of Geology & Geophysics, Vol.25.

Keyes, I.W. (1979). Ikamauius, a new genus of fossil sawshark (Order Selachii: Family Pristiophoridae) from the Cenozoic of New Zealand. New Zealand Journal of Geology & Geophysics, Vol.22, Number 1.

Welten, M., et al. (2015). Evolutionary origins and development of saw-teeth on the sawfish and sawshark rostrum (Elasmobranchii; Chondrichthyes). R.Soc.Open Sci., 12.

Share this post

Link to post
Share on other sites

Created August 4, 2016 due to length constraints.


Share this post

Link to post
Share on other sites

Updated September 11, 2016.



Share this post

Link to post
Share on other sites

Updated June 6, 2017.



Share this post

Link to post
Share on other sites

Updated October 9, 2017.



Share this post

Link to post
Share on other sites

Updated May 25, 2018.



Share this post

Link to post
Share on other sites

Create an account or sign in to comment

You need to be a member in order to leave a comment

Create an account

Sign up for a new account in our community. It's easy!

Register a new account

Sign in

Already have an account? Sign in here.

Sign In Now

Sign in to follow this  

  • Recently Browsing   0 members

    No registered users viewing this page.