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Fruitbat's Pdf Library - Class Asteroidea: Starfish

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These are a few of the pdf files (and a few Microsoft Word documents) that I've accumulated in my web browsing.

MOST of these are hyperlinked to their source.

If you want one that is not hyperlinked or if the link isn't working, e-mail me at joegallo1954@gmail.com and I'll be happy to send it to you.


Please note that this list will be updated continuously as I find more available resources. All of these files are freely available on the Internet so there should be no copyright issues.



Articles with author names in RED are new additions since May 11, 2018.


Phylum Echinodermata

Class Asteroidea - Starfish


Asteroidea - Africa/Middle East


Blake, D.B., L. Angiolini and A. Tintori (2014). Omanaster imbricatus (Echinodermata, Asteroidea), A New Genus and Species from the Sakmarian (Lower Permian) Saiwan Formation of Oman, Arabian Peninsula. Revista Italiana di Paleontologia e Stratigrafia, Vol.120, Number 3.

Gale, A. and L. Villier (2013). Mass Mortality of an Asteriid Starfish (Forcipulatida, Asteroidea, Echinodermata) from the Late Maastrichtian (Late Cretaceous) of Morocco. Palaeontology, Vol.56, Part 3.

Ghaderi, A. and L. Villier (2013). First record of the Late Cretaceous starfish Metopaster parkinsoni (Forbes, 1848) in Iran. Annales de Paleontologie, 99.


Asteroidea - Antarctica


Williams, M., et al. (2006). Late Miocene Asterozoans (Echinodermata) in the James Ross Island Volcanic Group. Antarctic Science, 18(1).


Asteroidea - Asia/Malaysia/Pacific Islands


Kato, M. and T. Oji (2013). A new species of Doraster (Echinodermata: Asteroidea) from the lower Miocene of central Japan: implications for its enigmatic paleobiogeography. Paleontological Research, Vol.17, Number 4.

Srivastava, D.K., et al. (2010). Record of Advenaster  Hess, 1955 (Asteroidea) from the Bathonian Patcham Formation of Kala Jhar in Habo Dome, Kachchh Basin, India. Journal of the Palaeontological Society of India, Vol.55(1).


Asteroidea - Australia/New Zealand


Eagle, M.K. (1999). A new Early Miocene Pseudarchaster (Asteroidea: Echinodermata) from New Zealand. New Zealand Journal of Geology & Geophysics, Vol.42.

Fell, H.B. (1954). New Zealand Fossil Asterozoa 3. Odontaster priscus sp.nov. from the Jurassic. Transactions of the Royal Society of New Zealand, Vol.82, Part 3.

Kesling, R.V. (1969). Three Permian Starfish from Western Australia and Their Bearing on Revision of the Asteroidea. Contributions from the Museum of Paleontology - The University of Michigan, Vol.22, Number 25.

Asteroidea - Europe (including Greenland and Siberia)


Blake, D.B. (2009). Re-evaluation of the Devonian family Helianthasteridae Gregory, 1899 (Asteroidea: Echinodermata). Palaontol. Z., 83.

Blake, D.B. (1996). Redescription and Interpretation of the Asteroid Species Tropidaster pectinatus from the Jurassic of England. Palaeontology, Vol.39, Part 1.

Blake, D.B. (1993). A New Asteroid Genus from the Jurassic of England and its Functional Significance. Palaeontology, Vol.36, Part 1.

Blake, D.B. and S. Rozhnov (2007). Aspects of life mode among Ordovician asteroids: Implications of new specimens from Baltica. Acta Palaeontologica Polonica, 52(3).

Blake, D.B. and J.W.M. Jagt (2005). New latest Cretaceous and earliest Paleogene asteroids (Echinodermata) from The Netherlands and Denmark and their palaeobiological significance. Bulletin De L'Institut Royal Des Sciences Naturelles De Belgique, 75.

Blake, D.B., A. Tintori and T. Kolar-Jurkovsek (2017). New Triassic Asteroidea (Echinodermata) Specimens and Their Evolutionary Significance. Revista Italiana di Paleontologia e Stratigrafia, Vol.123(2).

Blake, D.B., A. Tintori and H. Hagdorn (2000). A New, Early Crown-Group Asteroid (Echinodermata) from the Norian (Triassic) of Northern Italy. Revista Italiana di Paleontologia e Stratigrafia, Vol.106, Number 2.

Botting, J.P., et al. (2011). A new species of Siluraster (Echinodermata: Asteroidea) from the Late Ordovician of North Wales. Geological Journal, 46.

Gale, A.S. (2010). Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Department du Jura, France. Swiss J.Paleontol.

Hansen, T., D.L. Bruton, and S.L. Jakobsen (2005). Starfish from the Ordovician of the Oslo Region, Norway. Norwegian Journal of Geology, Vol.85, 3.

Herringshaw, L.G., M.P. Smith and A.T. Thomas (2007). Evolutionary and ecological significance of Lepidaster grayi, the earliest multiradiate starfish. Zoological Journal of the Linnean Society, 150.

Herringshaw, L.G., A.T. Thomas and M.P. Smith (2007). Starfish Diversity in the Wenlock of England. Palaeontology, Vol.50, Part 5.

Jagt, J.W.M. (1991). Early Miocene Luidiid Asteroids (Echinodermata, Asteroidea) from Winterswijk-Miste (The Netherlands). Contr.Tert.Quatern.Geol., 28(1).

Jagt, J.W.M. and V. Codrea (2010). A goniasterid starfish (Echinodermata, Asteroidea) preserved in a mid-Miocene rhyolitic ignimbrite, northwest Romania. Acta Geologica Polonica, Vol.60, Number 2.
Jagt, J.W.M., R.H.B. Fraaije and B.W.M. van Bakel (2009). A Late Miocene astropectinid (Echinodermata, Asteroidea) and associated ichnofossils from Liessel, province of Noord-Brabant, the Netherlands. Netherlands Journal of Geosciences, 88-2.

Neumann, C. (2000). Evidence of predation on Cretaceous sea stars from north-west Germany. Lethaia, Vol.33.

Neumann, C. and J.W.M. Jagt (2011). Pentasteria? splendida, a new Early Cretaceous astropectinid starfish from northern Germany. Swiss J.Palaeontol., 130.

Niebuhr, B. and E. Seibertz (2018). Comptoniaster michaelisi nom.nov. (Asteroidea, Goniasteridae): Revision of a starfish species from the lower Upper Cretaceous of central Europe previously described as Pentagonaster semilunatus and Asterias schulzii. Cretaceous Research, 87.

Repetto, G. and E. Bicchi (2013). Fossil starfishes (Echinodermata, Asteriidae) and paleontological analysis of the Pliocene of Cherasco, Piedmont region (NW Italy). Biodiversity Journal, 4(2).

Sutton, M.D., et al. (2005). A starfish with three-dimensionally preserved soft parts from the Silurian of England. Proc.R.Soc.B, 272.

Tinn, O. and L. Ainsaar (2014). Asterozoan pedicellariae and ossicles revealed from the Middle Ordovician of Baltica. Acta Palaeontologica Polonica, 59(2).

Villier, L. (2010). Asteroids from Barremian calciturbidites of the Serre de Bleyton (Drôme, SE France). Ann.Naturhist.Mus. Wien, Series A, 112.

Villier, L. (2008). Sea star ossicles from the Callovian black clays of the Lukow area, eastern Poland. N.Jb.Geol.Palaont.Abh., Vol.247/2.0.

Villier, L., S. Charbonnier and B. Riou (2009). Sea Stars from the Middle Jurassic Lagerstätte of La Voulte-Sur-Rhone (Ardeche, France). J.Paleont., 83(3).

Villier, L., M. Kutscher and C.L. Mah (2004). Systematics and palaeoecology of middle Toarcian Asteroidea (Echinodermata) from the "Seuil du Poitou", Western France. Geobios, 37.

Villier, L., et al. (2004). A preliminary phylogeny of the Pterasteridae (Echinodermata, Asteroidea) and the first fossil record: Late Cretaceous of Germany and Belgium. Palaontologische Zeitschrift, 78(2).

Zitt, J. (2005). Geinitzaster gen.n. (Asteroidea, Echinodermata) from Upper Cenomanian strata of the Bohemian Cretaceous Basin. Geologica Carpathica, 56, 4.


Asteroidea - North America


Blake, D.B. (2017). Two new Carboniferous Asteroidea (Echinodermata) of the family Urasterellidae. N.Jb.Geol.Palaont. Abh., 284/1.

Blake, D.B. (2010). Comptoniaster adamsi nov.sp. (Echinodermata, Asteroidea) from the middle Cretaceous of Texas and its phylogenetic position. Geobios, 43.

Blake, D.B. (2002). Compsaster formosus WORTHEN & MILLER (Asteroidea; Echinodermata): A Carboniferous homeomorph of the post-Paleozoic Asteriidae. Palaontol. Z., 76(2)

Blake, D.B. and R.W. Portell (2011). Kionaster petersonae, n.gen. and sp. (Asteroidea), the first fossil occurrence of the Asterodiscididae, from the Miocene of Florida. Swiss J. Palaeontol., 130.

Blake, D.B. and R.W. Portell (2009). Implications for the Study of Fossil Asteroidea (Echinodermata) of New Genera and Species from the Eocene of Florida. J.Paleont., 83(4).

Blake, D.B. and T.E. Guensburg (2005). Implications of a New Early Ordovician Asteroid (Echinodermata) for the Phylogeny of Asterozoans. J.Paleont., 79(2).

Blake, D.B. and T.E. Guensburg (1994). Predation by the Ordovician asteroid Promopalaeaster on a pelecypod. Lethaia, Vol.27.

Blake, D.B. and J.-P. Zonneveld (2004). Carniaster orchardi New Genus and Species (Echinodermata: Asteroidea), the First Triassic Asteroid from the Western Hemisphere. J.Paleont., 78(4).

Blake, D.B. and B.S. Kues (2002). Homeomorphy in the Asteroidea (Echinodermata); A New Late Cretaceous Genus and Species from Colorado. J.Paleont., 76(6).

Blake, D.B., W.K. Halligan and N.L. Larson (2018). A new species of the asteroid genus Betelgeusia (Echinodermata) from methane seep settings, Late Cretaceous of South Dakota. Journal of Paleontology.

Blake, D.B., et al. (2007). A New, Phylogenetically Significant Early Ordovician Asteroid (Echinodermata). J.Paleont., 81(6).

Gunderson, J.A. Asteroid Fossils from the Upper Jurassic of South-Central Montana. MBMG Open-File Report 660.

Kesling, R.V. (1982). Arkonaster, A New Multi-Armed Starfish from the Middle Devonian Arkona Shale of Ontario. Contributions from the Museum of Paleontology - The University of Michigan, Vol.26, Number 6.

Kesling, R.V. (1971). Michiganaster inexpectatus, a New Many-Armed Starfish from the Middle Devonian Rogers City Limestone of Michigan. Contributions from the Museum of Paleontology - The University of Michigan, Vol.23, Number 16.

Kesling, R.V. (1969). Silicaster, A New Genus of Devonian Starfish. Contributions from the Museum of Paleontology - The University of Michigan, Vol.22, Number 19.

Kesling, R.V. (1967). Neopalaeaster enigmaticus, New Starfish from Upper Mississippian Paint Creek Formation in Illinois. Contributions from the Museum of Paleontology - The University of Michigan, Vol.XXI, Number 3.

Kesling, R.V. and H.L. Strimple (1966). Calliasterella americana, A New Starfish from the Pennsylvanian of Illinois.Journal of Paleontology, Vol.40, Number 5.
Kesling, R.V. and J.D. Wright (1965). Two New Middle Devonian Species of the Starfish Devonaster from Southwestern Ontario.Contributions from the Museum of Paleontology - The University of Michigan, Vol.XX, Number 2.

Schuchert, C. (1915). Revision of Paleozoic Stelleroidea with Special Reference to North American Asteroidea. Smithsonian Institution, Bulletin 88. (400 pages, 61.1 MB download)

Williams, S.R. (1914). A Starfish Found in the Whitewater Division of the Richmond on Blue Creek, Adams County, Ohio. The Ohio Naturalist, Vol.XIV, Number 3.

Asteroidea - South America/Central America/Caribbean


Fernández, D.E., et al. (2014). An Early Cretaceous astropectinid (Echinodermata, Asteroidea) from Patagonia (Argentina): A new species and the oldest record of the family for the Southern Hemisphere. Andean Geology, 41(1).

Jagt, J.W.M., et al. (2013). A starfish bed in the Middle Miocene Grand Bay Formation of Carriacou, The Grenadines (West Indies). Geol.Mag., 151(3).

General Asteroidea


Blake, D.B. (2000). The Class Asteroidea (Echinodermata): Fossils and the Base of the Crown Group. Amer.Zool., 40.

Blake, D.B. and F.H.C. Hotchkiss (2004). Recognition of the Asteroid (Echinodermata) Crown Group: Implications of the Ventral Skeleton. J.Paleont., 78(2).

Gale, A.S. (2011). The Phylogeny of Post-Palaeozoic Asteroidea (Echinodermata, Neoasteroidea). The Palaeontological Society, London, Special Papers in Palaeontology Number 85. (119 pages)

Herringshaw, L.G., M.P. Smith and A.T. Thomas (2007). Evolutionary and ecological significance of Lepidaster grayi, the earliest multiradiate starfish. Zoological Journal of the Linnean Society, 150.

Mah, C. (2005). The phylogeny of Iconaster and Glyphodiscus (Echinodermata, Asteroidea, Valvatida, Goniasteridae) with descriptions of four new species. Zoosystema, 27(1).

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Updated May 11, 2018.



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