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Thecosmilia Trichitoma

Oldest known footprints in Grand Canyon found

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The abstracts related to this trackway are:


Rowland, Stephen M., 2018, Trackway of a Sideways-Walking Basal 

Tetrapod in the Pennsylvanian Manakacha Formation in Grand Canyon

 National Park. Abstracts of 78th Annual Meeting, Albuquerque 

Convention Center Albuquerque, New Mexico, U.S.A. October 

17–20, 2018. Society of Vertebrate Paleontology. p, 206 





Rowland, Stephen M., 2017, Trackway of a Sideways-Walking Pelycosaur

in the Carboniferous Manakacha Formation in Grand Canyon National Park.

Geological Society of America Abstracts with Programs. Vol. 49, No. 6, 






Paul H.

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