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I've been looking at my specimens of Acrocoelites trisulculosus from the Toarcian Jet Rock (Mulgrave Shale Member = Falciferum Zone) of the north Yorkshire coast. This is an anoxic mudstone deposited during a prominent worldwide Ocean Anoxic Event (OAE) and, as might be expected, the preservation is very good. A number of them (7 so far) have a thin pyrite layer around the apex. This shows obvious lineation in all of them, mostly oblique to the axis of the rostrum. As pyrite is often associated with soft tissue decay, I strongly suspect that this is preserving musc