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These are a few of the pdf files (and a few Microsoft Word documents) that I've accumulated in my web browsing.

MOST of these are hyperlinked to their source.

If you want one that is not hyperlinked or if the link isn't working, e-mail me at joegallo1954@gmail.com and I'll be happy to send it to you.


Please note that this list will be updated continuously as I find more available resources. All of these files are freely available on the Internet so there should be no copyright issues.


Articles with author names in RED are new additions since December 28, 2016.


Order Eulipotyphla (= 'Insectivora') - Hedgehogs, Shrews, Moles and Their Relatives


Family Dimylidae


van den Hoek Ostende, L.W. and O. Fejfar (2015). All time high: Dimylidae (Eulipotyphla, Mammalia) diversity in the early Miocene locality of Ahnikov 1 (Czech Republic, MN 3). Palaeobiodiversity and Palaeoenvironments, 95(3).

Ziegler, R. (2005). Erinaceidae and Dimylidae (Lipotyphla) from the Upper Middle Miocene of South Germany. Senckenbergiana lethaea, 85(1).


Family Erinaceidae - Hedgehogs and Gymnures


Subfamily Amphilemurinae


Smith, T., et al. (2002). A New Species of Macrocranion (Mammalia, Lipotyphla) from the Earliest Eocene of North America and its Biogeographic Implications. Contributions from the Museum of Paleontology - The University of Michigan, Vol.30, Number 14.


Subfamily Erinaceinae - Hedgehogs


Bi, S.-D. (1999). Metexallerix from the Early Miocene of North Junggar Basin, Xinjiang Uygur Autonomous Region, China. Vertebrata PalAsiatica, 37(2).

Engesser, B. and X.-L. Jiang (2011). Odontological and Craniological Comparisons of the Recent Hedgehog Neotetracus with Hylomys and Neohylomys (Erinaceidae, Insectivora, Mammalia). Vertebrata PalAsiatica, 49(4).

Lopatin, A.V. (2003). A New Genus of the Erinaceidae (Insectivora, Mammalia) from the Oligocene of Mongolia. Paleontological Journal, Vol.37, Number 6.

Matthew, W.D. (1903). A Fossil Hedgehog from the American Oligocene. Bulletin American Museum of Natural History, Vol.XIX, Article VII.

Meng, J., et al. (1999). The Petrosal Morphology of a Late Oligocene Erinaceid from North Junggar Basin. Vertebrata PalAsiatica, 37(4).

Schiebout, J.A. (1996). A Miocene Hedgehog (Mammalia: Erinaceidae) from Fort Polk in Western Louisiana. Occasional Papers of the Museum of Natural Science - Louisiana State University, Number 70.


Subfamily Galericinae - Gymnures  and Moonrats


Jenkins, P.D. and M.F. Robinson (2002). Another variation on the gymnure theme: description of a new species of Hylomys (Lipotyphla, Erinaceidae, Galericinae). Bull.nat.Hist.Mus.Lond. (Zool.), 68(1).

Klietmann, J., et al. (2014). Amphiperatherium and Erinaceidae of Petersbuch 28. Bulletin of Geosciences, 89(1).

Lopatin, A.V. (2004). A New Genus of Galericinae (Erinaceidae, Insectivora, Mammalia) from the Middle Eocene of Mongolia. Paleontological Journal, Vol.38, Number 3.

Masini, F. and F. Fanfani (2013). Apulogalerix pusillus nov.gen., nov.sp., the small-sized Galericinae (Erinaceidae, Mammalia) from the "Terre Rosse" fissure filling of the Gargano (Foggia, South-Eastern Italy). Geobios, 46.

Prieto, J., L.W. van den Hoek Ostende and M. Bohme (2011). Reappearance of Galerix (Erinaceomorpha, Mammalia) at the Middle to Late Miocene transition in South Germany: biostratigraphic and palaeoecologic implications. Contributions to Zoology, 80(3).

van den Hoek Ostende, L.W. and C.S. Doukas (2003). Distribution and evolutionary history of the Early Miocene erinaceid Galerix symeonidisi Doukas, 1986. In: Distribution and Migration of Tertiary Mammals in Eurasia. A Volume in Honour of Hans de Bruijn. Reumer, J.W.F. and W. Wessels (eds.), DEINSEA 10.

Villier, B. and G. Carnevale (2013). A New Skeleton of the Giant Hedgehog Deinogalerix from the Miocene of Gargano, Southern Italy. Journal of Vertebrate Paleontology, 33(4).


Subfamily 'Uncertain'


Gawne, C.E. (1968). The Genus Proterix (Insectivora, Erinaceidae) from the Upper Oligocene of North America. American Museum Novitates, Number 2315.


General Erinaceidae


Bannikova, A.A., et al. (2014). Contrasting evolutionary history of of hedgehogs and gymnures (Mammalia: Erinaceomorpha) as inferred from a multigene study. Zoological Journal of the Linnean Society, 112.

Frost, D.R., W.C. Wozencraft and R.S. Hoffmann (1991). Phylogenetic Relationships of Hedgehogs and Gymnures (Mammalia: Insectivora: Erinaceidae). Smithsonian Contributions to Zoology, Number 518.

He, K., et al. (2012). An Estimation of Erinaceidae Phylogeny: A Combined Analysis Approach. PLoS ONE, 7(6).

Lopatin, A.V. (2005). Late Paleogene Erinaceidae (Insectivora, Mammalia) from the Ergilin Dzo Locality, Mongolia. Paleontological Journal, Vol.39, Number 1.

Novacek, M.J., T.M. Bown and D. Schankler (1985). On the Classification of Early Tertiary Erinaceomorpha (Insectivora, Mammalia). American Museum Novitates, Number 2813.

Prieto, J. and M. Rummel (2009). Erinaceidae (Mammalia, Erinaceomorpha) from the Middle Miocene fissure filling Petersbuch 68 (Southern Germany). Zitteliana, A48/49.

Ziegler, R. (2005). Erinaceidae and Dimylidae (Lipotyphla) from the Upper Middle Miocene of South Germany. Senckenbergiana lethaea, 85(1).


Family Heterosoricidae


Furio, M., J. Prieto and L.W. vanden Hoek Ostende (2015). Three million years of "Terror Shrew" (Dinosorex, Eulipotyphla, Mammalia) in the Miocene of the Valles-Penedes Basin (Barcelona, Spain). C.R. Palevol, xxx. (Article in Press)


Family Palaeoryctidae


Bloch, J.I., R. Secord and P.D. Gingerich (2004). Systematics and Phylogeny of Late Paleocene and Early Eocene Palaeoryctinae (Mammalia, Insectivora) from the Clarks Fork and Bighorn Basins, Wyoming. Contributions from the Museum of Paleontology - The University of Michigan, Vol.31, Number 5.

Gingerich, P.D. (1982). Aaptoryctes (Palaeoryctidae) and Thelysia (Palaeoryctidae?): New Insectivorous Mammals from the Late Paleocene and Early Eocene of Western North America. Contributions from the Museum of Paleontology - The University of Michigan, Vol.26, Number 3.

Kielan-Jaworowska, Z. (1975). Preliminary Description of Two New Eutherian Genera from the Late Cretaceous of Mongolia. Palaeontologia Polonica, Number 33.

Lopatin, A.V. and A.O. Averianov (2004). New Palaeoryctidae (Mammalia) from the Eocene of Kyrgyzstan and Mongolia. Paleontological Journal, Vol.38, Number 5.

Thewissen, J.G.M. and P.D. Gingerich (1989). Skull and Endocranial Cast of Eoryctes melanus, a New Palaeoryctid (Mammalia: Insectivora) from the Early Miocene of Western North America. Journal of Vertebrate Paleontology, 9(4).


Family Solenodontidae


Morgan, G.S. and J.A. Ottenwalder (1993). A New Extinct Species of Solenodon (Mammalia: Insectivora: Solenodontidae) from the Late Quaternary of Cuba. Annals of the Carnegie Museum, Vol.62, Number 2.

Morgan, G.S., C.E. Ray and O. Arredondo (1980). A Giant Extinct Insectivore from Cuba (Mammalia: Insectivora: Solenodontidae). Proc.Biol.Soc.Wash., 93(3).

Roca, A.L., et al. (2004). Mesozoic origin for West Indian insectivores. Nature, Vol.429.


Family Soricidae - Shrews


Subfamily Crocidurinae


Botka, D. and L. Meszaros (2015). Crocidura (Mammalia, Soricidae) remains from the late Early Pleistocene Somssich Hill 2 Locality (Villany Hills Southern Hungary). Fragmenta Palaeontologica Hungarica, Vol.32.

Geraads, D. (1993). Middle Pleistocene Crocidura (Mammalia, Insectivora) from Oulad Hamida I, Morocco, and their phylogenetic relationships. Proc.Kon.Ned.Akad.v. Wetensch., 96(3).

Hugueny, M., et al. (2015). Lartetium africanum (Lavocat, 1961)(Eulipotyphla: Soricidae) from Beni-Mellal (Morocco), the oldest African shrew: new descriptions, palaeoenvironment and comments on biochronological context. Palaeobiodiversity and Palaeoenvironments, 95(3).

Hutterer, R. and D. Kock (2002). Recent and ancient record of shrews from Syria, with notes on Crocidura katinka Bate, 1937 (Mammalia: Soricidae). Bonn zool.Beitr., Bd.50, H.3.

Meszaros, L. (2015). Palaeoecology of the Early Pleistocene Somssich Hill 2 locality (Hungary) based on Crocidura and Sorex (Mammalia, Soricidae) occurrences. Hantkeniana, 10.


Subfamily incertae sedis


Tong, Y. and J. Wang (1993). A New Soricomorph (Mammalia, Insectivora) from the Early Eocene of Wutu Basin, Shandong, China. Vertebrata PalAsiatica, 31(1).


Subfamily Myosoricinae


Subfamily Soricinae


Botka, D. and L. Meszaros (2014). Beremendia (Mammalia, Soricidae) remains from the late Early Pleistocene Somssich Hill 2 Locality (Southern Hungary) and their taxonomic, biostratigraphical, palaeogeological and palaeobiogeographical relations. Fragmenta Palaeontologica Hungarica, Vol.31.

Czaplewski, N.J., J.O. Farlow and A. Argast (2012). A Fossil Shrew (Mammalia, Soricidae) from the Pipe Creek Sinkhole (Late Neogene: Hemphillian), Indiana. Proceedings of the Indiana Academy of Science, 121(1).

Czaplewski, N.J., et al. (2015). Late Pleistocene Shrews and Bats (Mammalia: Soricomorpha and Chiroptera) from Terapa, A Neotropical-Nearctic Transitional Locality in Sonora, Mexico. The Southwestern Naturalist, 59(4).

Dokuchaev, N.E., N. Kohno and S.D. Ohdachi (2010). Reexamination of fossil shrews (Sorex spp.) from the Middle Pleistocene of Honshu Island, Japan. Mammal Study, 35.

Emslie, S.D. (2002). Fossil Shews (Insectivora: Soricidae) from the Late Pleistocene of Colorado. The Southwestern Naturalist, 47(1).

Fanfani, F. (1998). Macroneomys sp. (Soricidae, Mammalia) from Visogliano Shelter (Trieste, Northern Italy), a site of Middle Pleistocene man. Acta zoological cracoviensia, 41(1).

Harrison, D.L., S.A. Parfitt and A.J. Stuart (2006). Occurrence of Macroneomys brachygnathus Fejfar, 1966 in the British Middle Pleistocene, with a review of the status of Beremendia fissidens (Petenyi, 1864) in Britain (Mammalia, Lipotyphla, Soricidae). Acta zoologica cracoviensia, 49A(1-2).

Hibbard, C.W. and D. Jammot (1971). The Shrews of the WaKeeney Local Fauna, Lower Pliocene of Trego County, Kansas. Contributions from the Museum of Paleontology - The University of Kansas, Vol.23, Number 24.

Jin, C.-Z., C.-K. Sun and Y.Q. Zhang (2007). Lunanosorex (Insectivora, Mammalia) from the Pliocene of North China. Vertebrata PalAsiatica, 45(1).

Lopatin, A.V. and A.S. Tesakov (2004). The fossil shrew Cretasorex arkhangelskyi Nessov et Gureev, 1981 from Uzbekistan - the systematic position among Soricidae, taxonomic status, and geological age. Russian J.Theriol., 3(1).

Meszaros, L. (2015). Palaeoecology of the Early Pleistocene Somssich Hill 2 locality (Hungary) based on Crocidura and Sorex (Mammalia, Soricidae) occurrences. Hantkeniana, 10.

Meszaros, L. (2014). A possible taphonomical evidence for the palaeoecological role of the giant shrews (Mammalia, Soricidae) in the Carpathian Basin. Hankeniana, 9.

Meszaros, L. (1999). Uppermost Pleistocene shrews (Mammalia, Soricidae) from Vaskapu Cave (N-Hungary). Annales Universitates Scientiarum Budapestinensis, Sectio Geologica, 32.

Parfitt, S.A. and D.L. Harrison (2011). New material of the shrew Macroneomys Fejfar, 1966 (Mammalia, Soricomorpha, Soricidae) from the British early Middle Pleistocene, with comments on its palaeobiology and European range. Acta zoologica cracoviensia, 54A(1-2).

Polly, P.D. (2003). Paleophylogeography of Sorex araneus: Molar Shape as a Morphological Marker for Fossil Shrews. Mammalia, 68(2).

Rofes, J., et al. (2016). Biostratigraphy, palaeogeography and palaeoenvironmental significance of Sorex runtonensis Hinton, 1911 (Mammalia, Soricidae): First record from the Iberian Peninsula. Palaeogeography, Palaeoclimatology, Palaeoecology, 459.

Rofes, J., et al. (2012). Nesiotites rafelinensis sp.nov., the earliest shrew (Mammalia, Soricidae) from the Balearic Islands, Spain. Palaeontologia Electronica, Vol.15, Issue 1.

Rzebik-Kowalska, B. (2013). Sorex bifidus n.sp. and the rich insectivore mammal fauna (Erinaceomorpha, Soricomorpha, Mammalia) from the Early Pleistocene of Żabia Cave in Poland. Palaeontologia Electronica, Vol.16, Issue 2.


General Soricidae


Gustafson, E.P. (1985). Soricids (Mammalia, Insectivora) from the Blufftop Local Fauna, Blancan Ringold Formation of Central Washington, And the Correlation of Ringold Formation Faunas. Journal of Vertebrate Paleontology, 5(1).

Menwer-Barakat, R., et al. (2010). Soricidae (Soricomorpha, Mammalia) from the Pliocene of Tollo De Chiclana (Gaudix Basin, Southern Spain). Journal of Vertebrate Paleontology, 30(2).


Family Talpidae - Moles


Subfamily Scalopinae - New World Moles


Barnosky, A.D. (1982). Locomotion in Moles (Insectivora, Proscalopidae) from the Middle Tertiary of North America. Science, New Series, Vol.216, Issue 4542.

Lloyd, K.J. and J.J. Eberle (2008). A new talpid from the late Eocene of North America. Acta Palaeontologica Polonica, 53(3).

Piras, P., et al. (2015). Digging Adaptation in Insectivorous Subterranean Eutherians. The Enigma of Mesoscalops montanensis Unveiled by Geometric Morphometrics and Finite Element Analysis. Journal of Morphology.

Yates, T.L. (1978). The Systematics and Evolution of North American Moles (Insectivora: Talpidae). Ph.D. Dissertation - Texas Tech University.


Subfamily Talpinae - Old World Moles


Hutterer, R. (1995). Archaeodesmana Topachevski & Pashkov, the correct name for Dibolia Rumke, a genus of fossil water moles (Mammalia: Talpidae). Bonn Zool.Beitr. Vol.45, Numbers 3-4.

Klietmann, J., et al. (2015). A gap in digging: the Talpidae of Petersbuch 28 (Germany, Early Miocene). Palaontol.Z., 89 (Author's personal copy)

Minwer-Barakat, R., A. Garcia-Alix and M. Freudenthal (2008). Desmaninae (Talpidae, Mammalia) from the Pliocene of Tollo de Chiclana (Guadix Basin, Southern Spain). Geobios, 41.

Rzebik-Kowalska, B. (2014). Review of the Pliocene and Pleistocene Talpidae (Soricomorpha, Mammalia) of Poland. Palaeontologia Electronica, Vol.17, Issue 2.

Sansalone, G., T. Kotsakis and P. Piras (2016). Condylura (Mammalia, Talpidae) reloaded: New insights about the fossil relatives of the genus. Palaeontologia Electronica, 19.3.54A.

Sansalone, G., T. Kotsakis and P. Piras (2016). New systematic insights about Plio-Pleistocene moles from Poland. Acta Palaeontologica Polonica, 61(1).

Schwermann, A.H. and T. Martin (2012). A partial skeleton of Geotrypus antiquus (Talpidae, Mammalia) from the Late Oligocene of the Enspel fossillagerstatte in Germany. Palaontol.Z., 86.

Skoczeń, S. (1993). New records of Parascalops, Neurotrichus and Condylura (Talpinae, Insectivora) from the Pliocene of Poland. Acta Theriologica, 38(2).

Ziegler, R. (2012). Moles (Talpidae) from Early Oligocene karstic fissure fillings in Southern Germany. Geobios, 45.

Ziegler, R. (2003). Moles (Talpidae) from the late Middle Miocene of South Germany. Acta Palaeontologia Polonica, 48(4).


General Talpidae


Meier, P.S., et al. (2013). Evolution of bone compactness in extant and extinct moles (Talpidae): exploring humeral microstructure in small fossorial mammals. BMC Evolutionary Biology 13:55.

Schwermann, A.H. and R.S. Thompson (2015). Extraordinarily Preserved Talpids (Mammalia, Lipotyphla) and the Evolution of Fossoriality. Journal of Vertebrate Paleontology, e934828. (Thanks to doushantuo for pointing this one out!)


General Eulipotyphla

General Eulipotyphla - Africa/Middle East


Furió, M., J. van Dam and F. Kaya (2014). New insectivores (Lipotyphla, Mammalia) from the Late Miocene of the Sivas Basin, Central Anatolia. Bulletin of Geosciences, 89(1).

Stanhope, M.J., et al. (1998). Molecular evidence for multiple origins of Insectivora and for a new order of endemic African insectivore mammals. PNAS, Vol.95.


General Eulipotyphla - Asia/Malaysia/Pacific Islands


Jin, C.-Z., K. Yoshinari and T. Hiroyuki (1999). Pliocene and Early Pleistocene Insectivore and Rodent Faunas from Dajushan, Qipanshan, and Haimao in North China and the Reconstruction of the Faunal Succession from the Late Miocene to Middle Pleistocene. Journal of Geosciences, Osaka city University, Vol.42, Article 1.

Lopatin, A.V. (2006). Early Paleogene Insectivore Mammals of Asia and Establishment of the Major Groups of Insectivora. Paleontological Journal, Vol.40, Supp.3.

Sulimski, A. (1969). On Some Oligocene Insectivore Remains from Mongolia. Palaeontologica Polonica, Number 21.

van den Hoek Ostende, L.W., G. Van Der Berch and R.A. Due. First fossil insectivores from Flores. Hellenic Journal of Geosciences, Vol.41.


General Eulipotyphla - Europe (including Greenland and Siberia)


Doukas, C.S. and L.W. van den Hoek Ostende (2006). Insectivores (Erinaceomorpha, Soricomorpha: Mammalia) from Karydia and Komotini (Thrace, Greece; MN 4/5). Beitr.Palaont., 30.

Furio, M. and C. Angelone (2010). Insectivores (Erinaceidae, Soricidae, Talpidae; Mammalia) from the Pliocene of Capo Mannu D1 (Mandriola, central-western Sardinia, Italy). N.Jb.Geol.Palaont. Abh., 258/2.

Furio, M., et al. (2015). The insectivores (Soricidae, Erinaceidae; Eulipotyphla; Mammalia) from Cueva Victoria (Early Pleistocene, Murcia, Spain). N.Jb.Geol.Palaont. Abh., 275/2.

Popov, V.V. (2004). Late Pliocene Erinaceidae and Talpidae (Mammalia: Insectivora) from Varshets (North Bulgaria). Acta zoologica cracoviensia, 47(1-2).

Prieto, J. (2011). The Miocene insectivores and marsupial from Affalterbach (North Alpine Foreland Basin, Germany). Zitelliana A, 51.

Prieto, J., L.W. van den Hoek Ostende and J. Hír (2012). The Middle Miocene insectivores from Sámsonháza 3 (Hungary, Nógrád County): Biostratigraphical and palaeoenvironmental notes near to the Middle Miocene Cooling. Bulletin of Geosciences, 87(2).

Prieto, J., et al. (2015). The Middle Miocene insectivores from Hasznos (Hungary, Nograd County). Palaeobiodiversity and Palaeoenvironments, 95(3).

Prieto, J., et al. (2010). Insectivores and bat (Mammalia) from the late Middle Miocene of Gratkorn (Austria): biostratigraphic and ecologic implications. N.Jb.Geol.Palaont.Abh., 258/1.

Rzebik-Kowalska, B. (2005). Erinaceomorpha and Soricomorpha (Mammalia) from the Miocene of Belchatow, Poland. IV. Erinaceidae Fischer Von Waldheim, 1817 and Talpidae Fischer Von Waldheim, 1817. Acta zoologica cracoviensia, 48A(1-2).

Rzebik-Kowalska, B. and L.I. Rekovets (2016). New data on Eulipotyphla (Insectivora, Mammalia) from the Late Miocene to the Middle Pleistocene of Ukraine. Palaeontologia Electonica, 19.1.9A.

Rzebik-Kowalska, B. and L.I. Rekovets (2015). Recapitulation of data on Ukrainian fossil insectivore mammals (Eulipotyphla, Insectivora, Mammalia). Acta Zoologica Cracoviensia, 58(2).

Sabol, M. (2005). Middle Miocene assemblage of insectivores from Bonanza site near Devinska Nova Ves (Slovakia). Geologica Carpathica, 56(5).

Smith, R. (2004). Insectivores (Mammalia) from the earliest Oligocene (MP 21). of Belgium. Netherlands Journal of Geosciences, 83(3).

Sulimski, A. (1962). Supplementary Studies on the Insectivores from Weze 1 (Poland). Acta Palaeontolica Polonica, Vol.VII, Numbers 3-4.

van den Hoek Ostende, L.W. and O. Fejfar (2006). Erinaceidae and Talpidae (Erinaceomorpha, Soricomorpha, Mammalia) from the Lower Miocene of Merkur-Nord (Czech Republic, MN 3). Beitr.Palaont., 30.


General Eulipotyphla - North America


Bown, T.M. and D. Schankler (1982). A Review of the Proteutheria and Insectivora of the Willwood Formation (lower Eocene), Bighorn Basin, Wyoming. U.S. Geological Survey Bulletin 1523.

Kirkland, H. and S. Norris (1999). Two Late Pleistocene Fossils from the Washita Local Fauna. Proc.Okla.Acad.Sci., 79:97-98.

Krishtalka, L. (1975). Systematics and Relationships of Early Tertiary Lipotyphla (Mammalia, Insectivora) of North America. Ph.D. Dissertation, Texas Tech University.

Mou, Y. (2011). Shrews, lagomorphs and rodents (excluding Cricetidae) from the Pliocene Panace Formation, southeastern Nevada, USA. Palaeontologia Electronica, Vol.14, Issue 3.


General Eulipotyphla


van den Hoek Ostende, L.W., J.W.F. Reumer and C.S. Doukas. The nature of the fossil record of Neogene insectivores. Hellenic Journal of Geosciences, Vol.44.

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Updated June 21, 2012.


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Updated April 6, 2013.


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Updated July 4, 2014.


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Reorganized and updated - August 1, 2015.


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Updated July 12, 2016.


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