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  1. From the album: Vertebrates

    Prismatoolithus gebiensis Zhao & Li, 1993 Upper Cretaceous Maojiacun Formation Neixiang Henan PRC Length 10cm / 4" These eggs probably stem from Troodons
  2. An interesting article “proving” most dinosaurs were warm blooded and that Troodon may have been able to switch back and forth to cold bloodedness. https://www.discovermagazine.com/planet-earth/fossil-eggshells-suggest-all-dinosaurs-may-have-been-warm-blooded https://news.yale.edu/2020/02/14/were-dinosaurs-warm-blooded-their-eggshells-say-yes
  3. Seen on an auction site. I highly doubt its a Troodon clutch but I thought I'd throw it out to the forums and see if anyone can ID it. Thanks
  4. Hello! Over the weekend I made some new labels for my fossil collection and I was wondering what everyone thought of them. I have QR codes which link to the corresponding "prehistoric-wildlife.com" species page for more info, and I added in some basic I.D. info to the cards to not crowd them. I also attached numbers to the labels and the fossils, so that I don't need to keep the labels directly next to the fossils. Would love to know what you think, and if anyone wants more information/the template I created. Thanks! P.S. Two of my I.D.s I'm still not 100% on (deltadromeus and Pectinodon) and I don't want anyone to assume I've completely I.D.ed them. Thanks!
  5. PaleoNoel

    Troodon (Stenonychosaurus?) Tooth

    From the album: Judith River fm. Fossil Finds

    This Troodon tooth is one of my favorite fossils in my whole collection. I found this at a microsite in Northern MT in the summer of 2017.
  6. This is being sold as a troodon foot claw from the Judith river, Montana. It measures 36mm. How does it look and does it look like the sickle claw found on the inner toe of each foot? Thanks in advance.
  7. fossils-uk

    Madagascan tooth - croc or dinosaur

    Hi guys just recently picked this up from the st Marie show. Not alot of information except it came from Madagascar. I bought this in conjunction with a rappetosaurus tooth also from Madagascar that has the same patina and look. the question is what is it? Crocodile? or is it something a bit more interesting and dinosaurian? always interested in your advice and help. best wishes byron
  8. CBchiefski

    Advanced Dinosaur Egg Guide

    The Advanced Dinosaur Egg Guide Please share this with those who have egg questions. When possible, technical terms were avoided or defined. Every effort has been made to ensure accuracy, but it is always important to do your own research. This guide is merely a snapshot of information taken from many scientific publications. I am not an expert on eggs, rather I just love sharing what little I have learned over the years, what science has learned over the years. For an overview on how to spot a fossilized dinosaur egg and the sizes of eggs, see the basic guide: Somewhat outdated yet still a good overview of dinosaur reproduction and eggs, with a focus on Mongolia: What is so special about eggs? The amniotic egg is one of the most significant evolutionary adaptations as it allowed vertebrate life to permanently exist on land. Long before the dinosaurs and their modern descendants including the chicken, the egg came first. In fact, the better question to ask is “Which came first? The lizard or the egg?” Before the amniotic egg, amphibians and some fish were the only vertebrates able to even venture on land and only for rather short periods of time. A great deal of information has come from studying eggs. What we have learned is summarized as: From University of California Museum of Paleontology Egg Anatomy: Using the best known modern avian dinosaur, the chicken--scientifically Gallus gallus, let us go over the different parts of an egg: “(A) The generalized anatomy of an egg. (B) The chicken eggshell comprises three crystalline layers, including the mammillary layer, prismatic layer, and external layer. The cuticle layer overlying the calcareous eggshell is further divided to two layers, including a HAp inner layer and a proteinaceous outer layer. The shell membrane, namely membrane testacea, is also characterized by two layers. (C) SEM image of the cuticle on the surface of the Gallus eggshell, showing a patchy and cracked pattern. (D) SEM image of the radial section of the Gallus eggshell. The white arrow indicates the cuticle layer that lies on the calcitic eggshell.” From Yang et al. 2018 Fig. 1 Those were technical terms, so how about we simplify. The chicken egg has three distinct shell layers mainly made of calcite, then a soft membrane on the inside of that. What is known as egg whites are the albumen which surrounds the yellow yolk located near the center. The embryo develops within the albumen and is fed with nutrients stored in the yolk. The surface of eggshell is full of openings, tiny pores, and these allow for gas to pass through the shell. A developing embryo needs to breathe just like any animal. Additional information: http://www.ucmp.berkeley.edu/science/eggshell/eggshell1.php How to spot a fake egg: First, the best way to avoid fake eggs is to go and collect them yourself. Always make sure to follow the laws and have permission to collect. In the United States, typically a good way to follow the law is through collection on private land with expressed permission from the landowner. Views of paleontologists do range on private ownership of fossils with many not condoning or endorsing. I personally have little issue with it since amateur collectors have made countless important finds while prospecting for their personal collection. If you are going to buy, do everything possible to ensure the egg or any fossil was legally collected. Often with fake eggs everything seems too perfect. Eggs are delicate and easily crushed or damaged so if there are no signs of any damage or natural alterations be very wary. If the surface has ridges, check to see those ridges continue across a crack or break of the shell. Many fake eggs are mosaics made up of real eggshell fragments assembled together in an egg shape. These mosaics tend to not have the eggshell match on opposite sides of a crack. If you would like more information beyond what is provided or have an unanswered question, feel free to start a thread. If after reading, you want to purchase an egg then please ask the seller for the best pictures they can provide of that egg with something to show scale such as a ruler and start a thread. There are many on the forum who are happy help determine if an egg is in fact real. Just please, whether collecting or buying, make sure you know the laws and follow them. A few good threads on real vs fake eggs: http://www.thefossilforum.com/index.php?/topic/69391-examples-of-commonly-faked-dino-eggs/ http://www.thefossilforum.com/index.php?/topic/83533-red-flag-on-hadrosaur-egg/ http://www.thefossilforum.com/index.php?/topic/71462-beware-of-hadrosaur-eggs/ http://www.thefossilforum.com/index.php?/topic/79465-this-is-how-realistic-a-fakereplica-oviraptor-egg-looks/ How are eggshell and eggs classified? Many people try to name an egg to a specific dinosaur, usually incorrectly. With embryonic remains, however, an egg can be scientifically linked to a particular dinosaur (explained in the next section). Another accepted way for eggs to be linked is through a pregnant female, there are examples of females which died while carry eggs internally. Adults on top of a clutch can be used however only with caution. Eggs are given their own naming scheme just as animals have theirs. In normal taxonomy, we have species, genus, and family whereas eggs have an oospecies, oogenus, and oofamily. The term used for egg taxonomy is parataxonomy. Parataxonomy is used in place of traditional taxonomy when an actual animal or plant cannot be linked, for example--from a lack of data. In the case of Troodon formosus, its eggs are the oofamily Prismatoolithidae, oogenus Prismatoolithus, and oospecies levis. Parataxonomy is the same system used for trace fossils, such as footprints which are normally not linked to the dinosaur who made them. What is inside a fossilized egg? Is there a yolk? What about bones? Very rarely are embryonic bones found, typically eggs have been filled in with sediments. These then lithify (become rock) and so the inside of nearly all fossil eggs is rock that is similar, if not identical, to the surrounding rock. Eggshell is brittle by its nature and so often cracks, these cracks allow whatever sediments are surrounding to fill in the egg and, depending on how recent it was laid to said crack, allow the amniotic sac and other fluids to drain out. Here is a CT scan of some eggs I am working on. You can see how the surrounding rock is very similar to the inside of the eggs. In addition to looking for embryonic material, the scan gives us information on the infill, the true shape of the eggs, and reveals anything which could otherwise not be seen within them. Sometimes insects can be found near an egg, for example. Embryonic bones from the oviraptor Citipati, this embryo is curled within the egg. From Wikimedia Commons Importance of Embryonic bones: https://youtu.be/cubdagTiRHE?t=48 Embryonic remains are vital for an actual animal ID, so any chance of them being present must be investigated. If you have any tiny bones which can be seen inside an egg or directly near it, I would strongly encourage you to take the specimen to your nearest paleontology related museum or university. If it does have embryonic remains in or near, then the specimen is invaluable to science. The presence of those tiny remains allows for the next question to be asked. Do we know who laid this egg? Which particular dinosaur? Most likely no, there are some wonderful exceptions though. Several ootaxa (eggshell type) are known to the dinosaur genus or family they were laid by. Here are some examples of eggs and eggshell which were linked scientifically to a particular dinosaur from embryonic remains. Dinosaur or family and its known egg type, oogenus or oofamily. This list is not comprehensive as new discoveries and revisions are made every year. Allosaurus sp. known to Preprismatoolithus coloradensis. (This is debated) Beibeilong (Oviraptor) known to Elongatoolithidae. Citipati (Oviraptor) known to Elongatoolithidae. (See the picture above) Gobipipus (Avian) known to Gobioolithus minor. Heyuannia (Oviraptor) known to Elongatoolithidae. Hypacrosaurus (Hadrosaur) known to Spheroolithus oosp. Lourinhanosaurus (Theropod) known to cf. Preprismatoolithus. Maiasaura (Hadrosaur) known to Spheroolithus oosp. Oviraptorid known to Elongatoolithidae. Therizinosauroid (med to large theropod) known to Dendroolithidae. Titanosaur (Sauropod) known to Megaloolithus patagonicus. Troodon (small Theropod) known to Prismatoolithus levis. Generally, be wary of any claim that an egg was laid by a certain dinosaur! Additional information: http://www.ucmp.berkeley.edu/science/eggshell/eggshell3.php What groups of dinosaurs do we have eggs for? The vast majority of eggs are from non-avian theropods. This group includes dromaeosaurs (like Velociraptor), allosaurs, and tyrannosaurs. We also have eggs from Mesozoic aves (birds), hadrosaurs (duck-billed dinosaurs) and sauropods (long-necks). It is worth noting when we say that the majority of eggs are therapod we mean it. Around 61% of the eggs found globally are therapod and between 41-64% are maniraptorans (birds and their closest non-avian dinosaur relatives). For the others the numbers are much smaller: 7% are sauropods, 13% are ornithischians (hadrosaurs and relatives) with 19% still unknown and that is no yolk. Here is an example of a clutch from an oviraptor, elongated eggs are typical of many theropods: Pic from The Zuhl Museum On the non-dinosaur side of things, we also have eggs from turtles, crocodiles, lizards, and pterosaurs (flying reptiles). There are several groups of dinosaurs who have no egg representation in the fossil record yet. Despite many people trying to find them, there are still no ceratopsian (horned dinosaur) eggs. There are no ankylosaur (armored dinosaur) or stegosaur (spiked/plated dinosaur) eggs as of yet either. This could simply be due to bias in the fossil record but there also could be other factors. Perhaps, it is a case like the ichthyosaur (marine reptile), which gave live birth, unlike most reptiles that lay eggs. Most of us are familiar with the platypus in the mammalian world, which lay eggs despite being a mammal. Maybe some dinosaurs did not actually lay eggs. Now that would be an eggciting discovery! Below one can see how similar clutches are for two very different types of hadrosaurs. The above is a rather typical egg clutch for a hadrosaur with spherical shaped eggs. Some of these eggs had embryonic remains which allowed them to be identified to a dinosaur. In this case they were narrowed down to within the lambeosaurinae subfamily but sadly could not be narrowed further. Pic from Museum of the Rockies Clutch of another hadrosaur, the good mother Maiasaura. Again, the eggs are spherical and embryonic remains allowed the eggs to be linked with Maiasaura. Pic from Museum of the Rockies The great identification mistake: Now that it is abundantly clear the only way to link a dinosaur and an egg is with embryonic bone. Why is that? Surely there must be other ways to ID who an egg is from. Well, let me share the story of poor Oviraptor, who was wrongly accused of stealing eggs. When the first Oviraptor was discovered, the skeleton was not alone. Underneath it was a clutch of eggs. At the time there were no embryonic remains in these eggs, so it was assumed that the strange looking animal was, in fact, stealing the eggs from Protoceratops, hence the name oviraptor meaning “egg thief.” Later, not far from the original site, another nest was found, this time with an almost perfectly preserved embryo. The embryo was clearly of that of an Oviraptor to be eggs-act. So, with both discoveries, paleontologists determined that Oviraptor was actually a brooding dinosaur much like birds today. This story is an eggcellent example of science improving upon itself and the need to be careful with assumptions. Paleontology is an ever-changing field, which constantly works to improve our understanding of the prior natural world. A common incorrect identification nowadays is that of “Tarbosaurus eggs.” Tarbosaurus is very similar to Tyrannosaurus rex, however, it lived in Asia. Among the largest of eggs ever found, were two measuring 11 cm (4.3 in) wide and an amazing 60 cm (24 in) long. The elongated shape meant they were probably from a large theropod and so were thought to be from Tarbosaurus. Scientifically these eggs are the oogenus macroelongatoolithus. Based on detailed analysis, these eggs most likely are from a large oviraptor and not Tarbosaurus. Alright, so then how are eggs differentiated and how without embryonic bones would an egg likely be from an oviraptor? How are eggs distinguished from each other? We went over how to link a dinosaur to an egg, what about one egg to another or finding differences between eggs? Well, there are a few different ways, one is the surface of eggshell. Many eggs have different textures but surface texture can be eroded or altered so cannot be used alone. Thickness and porosity of eggshell can be measured and provide solid data points for comparisons. Two of the best techniques for examining eggshell are with the use of SEM and thin sections. A scanning electron microscope (SEM) is a very powerful microscope, which can view objects in eggstreme detail. Petrographic thin sections are tiny slices of a rock so thin that light can actually pass through it. Both SEM and thin sections allow for the tiny details of eggshell to be visible, meaning unique traits, variations, and similarities can all be seen. Below are two types of eggshell, how many differences can you spot? A thin section of hadrosaur eggshell, there is only a single continuous layer. Pic from University of Calgary A thin section of oviraptor eggshell, there are two distinct layers with the arrow showing the point where both meet. Pic from University of Calgary On thick eggshell, the cross-section view can often show many details otherwise too small to see. Below is Faveoolithus eggshell, which is large enough to show the internal structure of the shell itself. Pic from Montana State University, taken by P. Germano Naming: Dinosaur eggs, much like actual dinosaurs, are named following a convention with information in the name, and normally an honor to an individual or location where it was discovered. As already covered, naming uses a system of parataxonomy and with eggs, this is called ootaxonomy. Using the method covered above, similarities and differences of eggshell can be identified. Based on these similarities and differences, eggs can be grouped. Some of these groups are associated with a type of dinosaur. As already covered, from embryonic remains or other methods an animal can be linked and associated to its eggs. Sometimes eggs can be grouped based on similarities yet there are no ways to associate them with a dinosaur, so these are listed as unknown. An egg group being associated to a type of dinosaur does not mean all eggs within the group are exclusive to that single type of dinosaur. Some eggs were named prior to the naming convention being established or do not fit any of the known groups, as such these have a truly unique name. That said, most eggs fit one of the following: Name- dinosaurs associated Sphero- Hadrosaurs Ovalo- Unknown Faveo- Unknown (Could be sauropods) Megalo- Titanosaurs Dictyo- theropods Dendro- Therizinosaurs Elongato- Oviraptors Prismato- Troodontids Egg and dinosaur associations, from top to bottom, Elongato- with Oviraptors, Sphero- with Hadrosaurs, Prismato- with Troodontids, Dictyo- and similar eggs from unknown theropods. Pic from the Royal Tyrrell Museum What time periods do we have eggs from? Nearly every egg from the Mesozoic is from within the Late Cretaceous. One study found of 238 eggs examined, 225 were from the Late Cretaceous, 10 from the Early Cretaceous, 2 from during the Late Jurassic and a single egg from the early Jurassic. Since then more eggs have been found, yet the trend holds. A likely explanation for such massive bias would be the Late Cretaceous is more recent so eggs from then are more likely to be preserved and undergo less alteration. Did an egg hatch? The hatching question is a difficult one to answer scientifically with most egg specimens, of course, a nearly complete egg is likely unhatched. Much of the strength in eggs comes from their shape and this means once there is an opening in the shell that strength is lost. There are many ways for an egg to break, one of which is the baby breaking out, but many of the broken eggs we find may have yielded no baby. The term unhatched and failed are often used interchangeably but the term failed is preferred as “unhatched” which implies the egg was fertilized and had a real chance. It is possible and likely probable that no fertilization was the cause for many eggs to not hatch. An overview of the different ways an egg can be filled. From Mueller-Towe et al. (2002) Nest? For as rare as eggs are, finding an egg clutch within a sedimentary structure is many times rarer. There have been several sedimentary structures found around egg clutches, which were interpreted as nests. One of the most interesting of these is a “U” shaped structure which looks similar to a horseshoe, see the picture below. In the center of this “U” shaped structure was a clutch of Troodon eggs. It is possible many nests were constructed like modern bird nests, with sticks, straw, leaves and other such material. This material in nest building, unfortunately, means they would most likely not preserve. Possible nest structure for Troodon, tape measure equals 1m (39in) and the white plaster jacket is covering a clutch of Troodon eggs. Modified from Varricchio et al. 1997 How can we tell what happened to an egg and the nest? By studying modern nests, it was found eggshell fragments tend not to travel very far while remaining in large concentrations. This means when a large grouping of eggshell fragments are found, it is unlikely they have moved much. Modern eggshell fragments can be found in ratios of concave up vs concave down based on what happened to the nest. For example, if a nest had a predator come and eat eggs, the eggshell would be concave up vs down in a ratio of about 70:30, sometimes 65:35. Obviously, if the eggshell fragments are moved then ratios will not work, but again, where high concentrations of eggshell are found, there was little to no movement. The ratio technique is still in the early stages of being applied to nest from the Mesozoic so in time there may be more information. The Emu eggshell above is concave-up. Pic by P. Germano The Emu eggshell below is concave down. Pic by P. Germano In both pictures, different layers of the eggshell can be seen and such layering indicates the eggshell is from a theropod, in this particular case, an avian. Where in the world are dinosaur eggs found? Eggs are extremely rare and there are only a select number of places where they have been found so far. Eggshell fragments, on the other hand, are actually rather common and can be found in many formations. One main reason eggshell is relatively abundant compared to complete eggs is that a single egg when broken can become dozens of fragments. Geographically eggs so far were found in Argentina, Canada, China, Columbia, France, Great Britain, India, Kazakhstan, Mongolia, Peru, Portugal, Romania, South Korea, Spain, Switzerland, the United States, and Uruguay. Within Canada, eggs are exclusively found in Alberta. Within the USA, eggs have been found in Colorado, Idaho, Montana, New Mexico, South Dakota, Utah, and Wyoming. The vast majority of eggs are found in Asia. Additional information: http://www.ucmp.berkeley.edu/science/eggshell/eggshell4.php Did dinosaurs care for their young? It seems that many dinosaurs did in fact care for their young. Evidence for this has been found on multiple continents. There is still debate over the type and amount of care the parents may have provided. There are two major variations in care being debated, and these come down to whether the offspring were altricial or precocial. See the list of terms near the end of this guide for definitions. One possibility is that a group of adults would use cooperative breeding to care for a clutch, this is basically the village raising a child approach. With theropods, in particular Oviraptor, the presence of adults on eggs does support incubation and possibly even brooding. Hatchlings have been found within a nest and could have died there for many reasons, brood reduction and siblicide are both entirely possible. Given the diversity of dinosaurs, it is likely different dinosaurs provided varying levels of care for their young. Modern example showing a female crocodile providing care: Modern example of a spoonbill bird raising young: Some dinosaurs such as the sauropod titanosaurs, likely did not care for their young but rather used the same strategy as sea turtles. A large group of females would lay hundreds of eggs at once to overwhelm the predators and just by sheer numbers allowing some of the babies to live to adulthood. Are there any diseases or mutations of eggshells? Yes, we have paleopathologies found in eggshell. Paleo meaning ancient and pathology being the study of diseases, so paleopathology is the study of ancient diseases. One of the more common is where two or more layers of eggshell overlap in a way where the pores no longer pass through the entire shell, this reduces the amount of oxygen an embryo can receive. Too many of the pores being misaligned can be fatal. What color were eggs? One of the most recent breakthroughs in egg research is an ability to determine colors present within fossilized eggshell. Interestingly, from the eggs so far examined there seem to be many colors and patterns. With this being rather new to the field, not many eggs have been tested plus there is likely some error and bias. Even so, there are remarkable results. Some eggs were simple, just white. Some were speckled. Many were dull earthy colors, while others were green and blue. Given their close relationship, it is logical to assume dinosaur eggs could show any variations of what we see from either crocs or birds. Modern crocodiles have white eggs whereas modern bird eggs range in color and pattern. Interestingly, even within the same bird species there is a range in color, so it is entirely possible dinosaur eggs from the same species also vary in color. Three modern chicken eggs showing variation in colors and size. From Wikimedia Commons What is working with eggs like? Fieldwork: The basic process of removing eggs from the ground is very similar to that of removing fossilized bones. The approximate size of an egg is figured out and then the area around it is trenched until a plateau is formed. Next, a plaster jacket is made encasing the plateau. The bottom of this is removed until the whole thing can be “popped.” After which it is flipped and then is ready to be brought back to the museum. An egg at a new nesting site just after I uncovered it. Pic from the Two Medicine Dinosaur Center Jacketing an egg at Egg Mountain in Montana. Pics by D. O’Farrell. To find small fragments of eggshell and embryonic bones, removed rock is often sifted. Since they are so small—and also a rock surrounded by rocks—many times until sifted, the tiny bones or eggshell are not visible. Sifting for eggshell, here I am showing Paleontologist Barbie an example eggshell fragment. Pic from Coffeewithhallelujah After viewing the example fragment, my esteemed colleague Paleontologist Barbie was able to find an eggshell fragment. Can you find the piece of eggshell below? Pic from Coffeewithhallelujah Preparing and reconstructing an egg: Eggs tend to be more tedious and require more patience than normal prep work. Eggs are not that difficult to prepare, however, to an even greater extent than bones, they are very unforgiving. Reassembling a fossil bone after a mistake is not necessarily easy, however it is normally possible. The same often cannot be said for fossilized eggs. If you ever want to try and reconstruct a dino egg, just save the last chicken egg after cracking it and then try to reassemble. Remember, chickens are dinosaurs and their eggs make a decent modern analog to a classic theropod egg. Eggs in context- The Two Medicine Formation: To bring us all the way back to the beginning, what is the importance of studying eggs? Why bother? The primary geologic formation I have spent the last seven years working in is the Two Medicine and in terms of eggs, it is the most significant location in North America. One newly discovered nest I am fortunate enough to have an ongoing role in excavating and scientifically describing. From eggs and embryonic remains, the ecosystem of the Two Medicine is relatively well known compared to nearly every other formation. In terms of paleoecology, nesting sites show where adults felt safe and secure with enough food, water, and other resources. Within this formation was true evidence for parental care, particularly care in the form of nurture similar to birds. Behavior is nearly impossible to deduce from the limited fossil record, yet the care for young is strongly supported thanks to discoveries in the Two Med. Three dinosaurs from the formation have been linked to their eggs, Hypacrosaurus, Maiasaura, and Troodon. It may not seem impressive but three dinosaurs with embryonic remains is a truly remarkable find and incredibly rare. Even now, after over forty years of study, the Two Med continues to surprise with new nesting sites. Read about how the Two Medicine and Maiasaura was discovered: Additional information: http://www.ucmp.berkeley.edu/science/eggshell/eggshell_case1.php https://www.nps.gov/articles/mesozoic-egg-mountain-dawson-2014.htm https://serc.carleton.edu/research_education/mt_geoheritage/sites/augusta_choteau/paleontology.html http://www.georgialifetraces.com/2014/07/15/tracing-the-two-medicine/ http://www.georgialifetraces.com/2014/08/04/fossil-visions-in-the-two-medicine/ Hear me talk about my research on eggs and Troodon: Dinosaurs as living animals: Eggs allow us to see these animals as just that, animals. There is a reason many feel sad when seeing a baby dinosaur still in its egg, yet the same sadness tends to not be shown for adults. Why? The poor baby was deprived of an actual life and it is easy to relate. When covering a natural disaster, one goal of reporting is to humanize the story. In a similar way, when reporting on dinosaurs, it is important to try and do the same. Eggs allow us to come far closer to dinosaurs as true animals than I feel we ever will through bones alone. Eggs and reproduction give a window into the lives of these wonderful animals. When trying to describe what separates something living from an inanimate object, the ability to reproduce is used as a major criterion, therefore making it one of the most important aspects of dinosaurs to study in detail. Some Relevant Terms: These typically are used for modern birds and the classic theropods. Altricial: A developmental classification where at hatching, the offspring are relatively immobile, lack feathers or down, have closed eyes and are completely dependent on their parents for survival. Altricial birds include herons, hawks, woodpeckers, owls, and most passerine songbirds. Brood (n): The offspring of an animal which are hatched or cared for at one time. Brood (v): To sit on and keep warm. Brooding: To sit on and keep offspring warm when they cannot maintain their own body temperatures. Brood reduction: A reproductive strategy where the female lays more eggs than can be cared for and raised. The smallest and weakest of the brood typically starve or are killed by siblings. Clutch: Total number of eggs laid by a female in one nest attempt, often 3 or more. Conspecific: Of the same species. Cooperative breeding: Breeding system where non-parental adults assist other breeding pairs (usually their own parents) to rear offspring, instead of dispersing from the nest or breeding themselves. Incubation: The process by which parents keep eggs at the proper temperature to ensure normal embryonic development until hatching. In most cases, birds sit on eggs and transfer their body heat through a patch of skin known as the brood patch. In many species, only the female incubates; in other species, both males and females incubate. Less common is where only the male incubates. Precocial: Offspring are capable of a high degree of independent activity immediately after hatching. Precocial young typically can move about, have their eyes open and will be covered in down at hatching. They are generally able to walk away from the nest as soon as they have dried off. Siblicide: The death of a young animal usually as a result of fighting with siblings over food, common in years when food is in short supply. Further reading and information: https://www.amnh.org/our-research/paleontology/about-the-division/more/fossil-identification/dinosaur-eggs-fossil-identification http://www.ucmp.berkeley.edu/science/eggshell/index.php http://www.ucmp.berkeley.edu/science/eggshell/eggshell_hirsch.php http://www.ucmp.berkeley.edu/science/eggshell/eggshell5.php https://feederwatch.org/blog/raptors-make-good-neighbors-hummingbirds/ Images: University of California Museum of Paleontology: http://www.ucmp.berkeley.edu/ Yang et al. 2018: https://doi.org/10.7717/peerj.5144 Montana State University: http://www.montana.edu/ Two Medicine Dinosaur Center: http://www.tmdinosaurcenter.org/ Royal Tyrrell Museum: http://tyrrellmuseum.com/ Museum of the Rockies: https://museumoftherockies.org/ The Zuhl Museum: https://zuhlmuseum.nmsu.edu/ Dr. Tony Martin: http://www.georgialifetraces.com/ Mueller-Towe et al. 2002: https://www.researchgate.net/publication/260391508_Hatching_and_infilling_of_dinosaur_eggs_as_revealed_by_computed_tomography University of Calgary Hadrosaur eggshell: https://www.ucalgary.ca/drg/imagesort/00S000500 Oviraptor eggshell: https://www.ucalgary.ca/drg/imagesort/00S001300 Varricchio et al. 1997: https://www.researchgate.net/publication/232793785_Nest_and_egg_clutches_of_the_dinosaur_Troodon_formosus_and_the_evolution_of_avian_reproductive_traits Coffeewithhallelujah: http://coffeewithhallelujah.blogspot.com/2015/07/paleontologist-barbie-at-two-medicine.html Wikimedia Commons Citipati: https://en.wikipedia.org/wiki/Citipati Chicken eggs: https://en.wikipedia.org/wiki/Egg_as_food List of open access egg related papers: Thanks to the late Joe Gallo for this wonderful list. Disclaimer: For legal purposes, it should be noted links to an institution does not constitute endorsement by the respective institution and pictures are used here for educational purposes only. All rights belong to their respective owners. From the 2018 SVP meeting, my poster, which was a presentation on new dinosaur eggs. Pic from the Two Medicine Dinosaur Center Many thanks to J. Cozart and L. Murphy for writing some sections as well as edits. Thanks to D. Lawver, Ph.D. for reviewing the information presented. I especially would like to thank @Fossildude19 for assisting me and additionally thank these members for input and suggestions: @Troodon . @Seguidora-de-Isis . @HamptonsDoc . @-Andy- Eric P.
  9. I thought I would share this amazing articulated Troodon skeleton recently found by a respected fossil team. It's currently on the market so hopefully it ends up in a nice museum so everyone can enjoy it.
  10. I thought I’d post some of my favorite claws from my collection. I’m curious what people think about my ID on one of them and I have no idea what the last one is. ID help with that one would be great! (All measurements are straight line) Spinosaurus hand claw 4 1/2” Kem Kem Beds, Morocco Repairs, but I see no restoration Acheroraptor Temerytorum foot walking claw 1 7/8” Hell Creek Carter County, Montana No repair or restoration (at first). However, the tip broke off during molding and it was lost. 1/16” restoration done to the tip now Two Acheroraptor killing claws. The larger one is 3 1/8” and the smaller is 1 9/16” Hell Creek Powder River County, Montana Large claw has restoration to the top 1/4” of the articulation end and 3/4” to the tip Smaller claw has restoration on 5/8” of the tip Same claw as above, with size perspective Other side... Acheroraptor Temertyorum digit I hallux claw 7/8” Hell Creek Slope County, North Dakota No repair or restoration Acheroraptor Temertyorum hand claw 1 1/4” Hell Creek Powder River County, Montana Restoration to 1/2” of the tip I originally thought this this was a Pachycephalosaurus claw, but Troodon’s posting on TFF makes me now believe it’s Thescelosaurus Hell Creek Powder River County, Montana No repair or restoration Side view.. Troodontid walking foot claw 1 1/16” Hell Creek Wibeaux County, Montana Looks like restoration to 1/4” of the tip Possible Microraptorine hand claw 5/8” Hell Creek Carter County, Montana Small amount of restoration to the top of the articulating end and 1/8” of the tip Same claw... Microraptorine killing claw. Related to Hesperonychus sp. 7/8” Hell Creek South Dakota Restoration to 3/16” of the tip Same claw for size comparison... Curious what people think of this one.. I believe it to be a Troodontid killing claw 3/4” Two Medicine Formation Unfortunately, no locality info on it Looks like the tip was glued back on, but no restoration Other side... Now I have no clue what this could be and I’d love some help. I bought it as a new collector awhile ago with very little knowledge. It was sold as a baby Anzu foot claw and the seller said Black Hills Museum ID’d it. I think it’s actually mammalian. Any thoughts? Size comparison Articulating end (sorry for the poor pics)
  11. Darko

    Troodon

    This 2019 is getting better and better ! Here is my new drawing,its Troodon! Hope you like it. Darko
  12. Hey hi All, Does anybody know where Frank went? @Troodon Hopping You are doing alright!!! Best, Tony
  13. I saw these teeth for sale and the Id's look squirrely but I'm not an expert so look for a positive ID. All from Hell Creek Formation The seller identifies this as a Troodon tooth 1/4" looks more like a Pectinodon The others are being sold as Dromaeosaurus/Raptor All are 1/2" or less. They look like juvie Nanotyrannus
  14. I noticed a number of online suppliers offering Troodon teeth that actually belong to the genus Pectinodon. Thought it would be a good discussion item for a focused post since they can get confusing with the various morphologies of Pectinodon teeth. I'll use publications to illustrate my points and reflect all Campanian & Maastrichtian age teeth of North America. (Edit) A new paper published late in 2017 has turned this taxon upside down. I will try to reflect those changes here but it should not change alter how they are identified against Pectinodon teeth. Specimens from Dinosaur Park Formation previously assigned to Troodon formosus have been reassigned to a new taxon Latenivenatrix mcmasterae and others to a resurrected Stenonychosaurus inequalis. Troodon formosus is now considered invalid. At the very end of this page I will show my understanding of ID's by Formation. Canadian Journal of Earth Sciences, 2017, Vol. 54, No. 9 : pp. 919-935 Troodontids (Theropoda) from the Dinosaur Park Formation, Alberta, with a description of a unique new taxon: implications for deinonychosaur diversity in North America Aaron J. van der Reest, Philip J. Currie https://doi.org/10.1139/cjes-2017-0031 (Edit 2) Troodon formosus is still considered valid, comments from publication: "Given that the latter had already been synonymized into the senior T. formosus and remained unused for 30 years, Troodon formosus remains the proper name for this taxon, exclusive of L. mcmasterae, and we continue to use it here.: https://www.nature.com/articles/s41598-018-30085-6 Troodontids - Standard morphology These teeth are recurved, laterally compressed and oval in cross section. The Denticles are large and pointed upward on the distal (posterior) carina and minute or absent on the mesial (anterior) carina. Premaxillary teeth have strong denticles on both edges (Figure 8.3 E) Denticles per mm is typically around 4 Maximum length about 9.8mm Figure Examples : Troodontids: Large Morphology The tooth is large about 50% greater and others and can reach 14mm long, recurved and round in cross section. (See figure below 8.3 # 20-21) These have been collected from the North Slope of Alaska. Their is nothing unique to these teeth to describe a new taxon. I've have seen one of these jumbo teeth come from Alberta. Troodon Alaska.pdf Pectinodon bakkeri : Comparison - Troodon and Pectinodon Pectinodon - left image #5 mesial denticles usually absent #11 posterior denticles very large and often rounded much smaller average length of 2.6mm Troodonitid right image #12 posterior denticles point to the tip #13 anterior denticles exist, can be large or absent much larger average length 4 mm but can reach 9.8 Scale bar 1mm Pectinodon teeth can be put into four categories: premaxillary, maxillary, anterior & posterior dentary and all look different. In Figure 9.5 A & B are Premaxillary, C & D Maxillary, E & F Anterior Dentary and G & H Posterior Dentary Premaxillary teeth: long and slender, mesial carina strongly convex and distal is straight. Distal Denticles are large, pointed to the tip and become smaller toward the base. Maxillary teeth are compressed and bladelike. They look like small Dromaeosaur teeth. Anterior denticles are irregular in size and very small (5-6 denticles/mm). Posterior denticles are 3 per/mm. Anterior teeth are leaf shaped with no serrations on the mesial carina. Denticles on the distal edge are 1.6/mm. Denticles are irregular in size. Please note scale bar at 5mm References used: 1) Vertebrate Microfossil Assemblages by Sankey and Baszio 2) Dinosaur Systematics by Ken Carpenter and P. Currie North American Troodontids: My best call I"m sure these will be updated with new research and discoveries. Belly River Group (Alberta) : Latenivenatrix mcmasterae and Stenonychosaurus inequalis (one cannot distinguish between isolated teeth of these two species ID: should be Troodontid indet.) Horseshoe Canyon Formation (Alberta) : Albertavenator curriei (multiple species may exist) Kaiparowits Formation (Utah) : Talos sampsoni Judith River Formation (Montana) : Troodon formosus Two Medicine Formation (Montana) : TMF Troodontid Hell Creek & Lance Formation (Montana, Wyoming, South & North Dakota) : cf Troodon formosus All others (Saskatchewan, Alaska, Colorado etc) : cf Troodon formosus Multiple Troodontids might exist in these other fauna similar to the Belly Group. Since skeletal remains are super rare we may never know if this is true. Identification to a family level "Troodontid indet." may be more appropriate but you can decide that.
  15. BonuFrailman

    Possible Troodontid tooth

    Hi, Saw what looks to be some kind of troodontid tooth online. Was wondering if anyone could ID it. Tooth is from Judith River formation and appears to measure less than an inch. Thanks.
  16. Recent, I came across several teeth from Kem Kem which intrigued me. At first thought, they reminded me of Troodon teeth, though their serrations were nowhere as big. One seller labelled it as theropoda indet. Another called it Masiakasaurus. I thought they might even be Abelisaurid teeth. What are your thoughts on these?
  17. Here are picks of my favorite dinosaur teeth that I have in my collection- "Troodon".
  18. http://www.timescolonist.com/hips-don-t-lie-unearthed-dinosaur-pelvic-bones-shake-up-family-tree-1.21715300 "As a result, Troodon formosus is no longer considered a valid species" (and @Troodon might have to change his name )
  19. -Andy-

    Protoceratops Egg?

    I just acquired what looks to be a very unusual egg. This single egg comes detached from a clutch of upright eggs in an antique shop in Singapore; owner would not reveal where it origins from. The egg is white, and covered in reddish-sand matrix that can be scrapped off with a needle. This egg measures 7.66 cm long (I lack measurement of the nest, owner wouldn't let me take more pics). My first thought were that these are Troodon eggs, as "upright eggs" are a characteristic of troodon eggs as far as I know. 1) Thomas(Tom) Kapitany said these looks like it came from China. He didn't personally give it an ID, but mentioned he had seen these identified as Protoceratops before. 2) Seth Sorenson said there's no way to confirm it as Troodon without skeletal material, but he said they look like small ceratopsian eggs to him. He also mentioned Protoceratops eggs have been found in standing clutches before. 3) Laogao, a fossil egg expert from China said this looks like Troodon eggs from Henan. 4) Dr. Kenneth Carpenter said these are most likely theropod eggs of the Elongatoolithus(egg oogenera) family. Without microscopic examination or confirmation of their origin, he could not give any further identification though. With such little information, I am aware there's no way to give a definitive ID of the dinosaur that laid these eggs. I'd still like to ask for your opinions though on what you believe this egg might be.
  20. AJ Plai

    Troodon tooth

    From the album: Dinosaur Fossils collection

    Troodon tooth Locality: Hell Creek Montana, USA Geological Age: Cretaceous Specimen Size: 0.45 cm
  21. Pterosaur

    Dinosaur Intelligence

    Hello everyone, I've been researching dinosaur intelligence, and have read about certain dinosaurs who were considered more intelligent than most others. From what I gather, even the cleverer dinosaurs were left in the dust by modern birds and mammals, yet dinosaurs were apparently very smart for reptiles... I've read that Ankylosaurids were exceedingly unintelligent, and that Troodon were brilliant for their time. The intellect of Ornithomimus is speculated by some to have been akin to that of an ostrich, and Tyrannosaurus Rex apparently had a fairly large section of its brain specifically devoted to strategy. I know animal intelligence (even human intelligence) is tough to measure, and I'd imagine that DINOSAUR intelligence can only be 100% more difficult. With that said, what I'm trying to ask is: How intelligent do you think dinosaurs were relative to today's birds, mammals and reptiles? How dim were massive sauropods and ankylosaurids compared to very dim modern animals? Was Troodon really just as smart as an opossum? Obviously this requires a lot of wild speculation, but it'd still be fun to hear your thoughts. I know there are people on this forum who specialize their interests and knowledge in particular dinosaurs, so please fire away. Thanks! Lauren
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