Desmostylus hesperus
There is no common name for this Miocene group of mammals which has unknown affinities, unknown origin, and unknown feeding habits. Desmostylians are restricted to the northern Pacific Rim, from Japan to Mexico. (Recent work rejects a questionable record from Florida.)
Desmostylians were probably amphibious, pony-sized with short, stout limbs. They seem to have been able to walk on the bottom, rather than swim, in shallow coastal areas.
Carbon isotope levels have been measured in the enamel of Middle Miocene Desmostylus teeth from California. Isotopic data suggest that Desmostylus was an aquatic herbivore that spent a considerable portion of its life foraging in estuarine and freshwater ecosystems.
Their skeleton has been described as horse-like. Their habitus may have been hippo-like.
These mammals had well-developed, forward-projecting canines and incisors in both upper and lower jaws. Cheek teeth apparently were replaced cyclically as in elephants.
The structure of the cheek teeth is unique among mammals. Each tooth is comprised of a cluster of stout dentin tubes covered with thick enamel. It is this structure that gave the group its name (desmos = chain, stylos = pillar).
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