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Partial Jurassic Neuroptera Insect with Beautiful Wing Preservation
Biotalker posted a topic in General Fossil Discussion
Here is my recent acquisition from Inner Mongolia of China in the Jiulongshan formation, more commonly and famously known as Daohugou, which dates to middle Jurassic around 165 mya. The part and counterpart partial insect fossil appears to be a neuropteran, perhaps related to the family of Grammolingiidae (a group of lacewings that include “Jurassic butterflies”, so named on account of their huge spotted wings). Only one of the wings and a small part of another were found in the fossil. The wing, however, is slightly more than two inches (5cm) long. The head and thorax but not the abdomen are intact. I am sharing these images because I think the fossilization is remarkable. Dense tissue like the head and thorax are harder to interpret than the thin wing. But wow, the details that can be seen under the scope are amazing! No ambiguity as we sometimes see with dragonfly fossils!- 8 replies
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There are additional unidentified conchostraca (clam shrimps, arthropods) on the slab. The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin, 2001 reported an Argon-argon dating age of 164 ±4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian); this opinion is now widely accepted. Taxonomy according to Fossilworks.org. Emended diagnosis for Choristopsychidae according to Qiao et al., 2013, p. 93: "Forewing broad oval or subtriangular, field between C and ScP comparatively broad; ScP well developed and forked twice, forming three long branches; RA unforked, one crossvein between ScP and RA and between RA and RP; RP and MA both with two branches; MP with five branches, and the MP4+5 forking basal to the MP2+3 forking; MP and CuA merged at the base; CuA strongly bent at its mid point; an oblique crossvein between CuA and CuP; a curved crossvein between the midpoint of CuA and MP5; CuP, 1A and 2A almost parallel. Hind wing, similar in shape to the forewing but slightly smaller, ScP short, forked twice, the second bifurcation coalesces with RA for a short distance; RP and MA both with two branches; MP with five branches, the stem of MP4+5 forked earlier than that of forewing, and with a crossvein to CuA; CuA almost straight. Head, oviform with big and oval compound eyes; antennae long and filiform; small chewing mouthpart. Thorax: prothorax smaller than mesothorax and metathorax. Legs: long and slender, all legs nearly of the same shape, but hind legs longer than fore legs and mid legs, and femora wider than tibia, and tibia longer than femora. Abdomen slender, tapering apically, about eleven segments and the female terminal segment with cercus." Choristopsyche asticta, line drawing of the holotype from Qiao et al., 2013, p. 100. Identified by Prof. A. Rasnitsyn (Russian Academy of Sciences) as Choristopsyche cf. asticta Qiao et al., 2013. Reference: Gao K Q, Shubin N. (2001): Late Jurassic salamanders from Northern China. Nature, 410: 574–577. Martynov, A. V. (1937): Liassic insects from Shurab and Kisyl-Kiya, Part I, Various orders except Blattodea and Coleoptera. Akademiya Nauk SSSR, Trudy Paleontologicheskogo Instituta 7: 1-178. Qiao, X., Shih, C. K., Petrulevičius, J. F., and Ren, D. (2013): Fossils from the Middle Jurassic of China shed light on morphology of Choristopsychidae (Insecta, Mecoptera). ZooKeys 318: 91-111. link
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The age of the Daohugou strata has been notoriously difficult to determine, and a number of studies have produced conflicting results. Gao and Shubin reported an Argon-argon dating age of 164+- 4 million years ago (Middle to Late Jurassic, Bathonian to Oxfordian), this oppinion is now widely accepted. Part and counterpart of the same fossil. There are additional unidentified conchostraca (clam shrimps, arthropods) on the slabs. Drawing of the holotype by Liang et al. 2012 Taxonomy according Fossilworks.org. Differential diagnosis (genus) according to Liang et al, 2012: "The new genus can be distinguished from other representatives of the family by the richly branched Sc. The new genus additionally differs from Liadoblattina Handlirsch, 1906 (Vršanský and Ansorge 2007) from Early Jurassic of Germany and England in having a larger body and reticulations present in the CuA-CuP area. Graciliblatta is also different from Raphidiomima and Cameloblatta in the elongated pronotum (the length-to-width ratio of the pronotum of the new genus is 1.5, those of Raphidiomima and Cameloblatta 1.2), and distinctive divided eyes located basally, which are undivided in the other genera, which originate from the Late Jurassic of Kazakhstan. The head and pronotum of Graciliblatta are extremely long and narrower than in Fortiblatta (Liang et al. 2009); vein R of the forewing is more arcuate and R1 has secondary branching not found in Fortiblatta, which is from the same locality and age as Graciliblatta. In addition, the body is smaller (length/width of head in the new genus is 2.25 vs. 1.9 in Fortiblatta; the pronotal length/width of the new genus is 1.5 vs. 1.2 in Fortiblatta). Coloration in the tip of wings is absent, but Graciliblatta has two dark stripes on pronotum, unlike Fortiblatta." Description (species) according to Liang et al., 2012: "Head and pronotum elongated. Forewing long and narrow. Sc richly branched, with 7–8 branches; R not reaching the tip of margin, slightly curved and basally with dark coloration. CuA and CuP slightly curved. Anal veins with tertiary branches. Diagonal fold present. Hind wing with simple Sc; R1 and Rs differentiated, and R1 secondarily branched." Prognathous and sharp mandibles of the species suggest it was a carnivore. Identified by oilshale. Reference: LIANG, J. H., HUANG, W. L. & REN, D. (2012) Graciliblatta bella gen. et sp. n. - A rare carnivorous cockroach (Insecta, Blattida, Raphidiomimidae) from the Middle Jurassic. Zootaxa 3449: 62–68. LIANG, J. H., SHIH, C. K. & REN D. (2018). New Jurassic predatory cockroaches (Blattaria: Raphidiomimidae) from Daohugou, China and Karatau, Kazakhstan. Alcheringa 42:101-109.
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Mecoptera (or scorpionfly) non det. from Nei Mongol, China.
oilshale posted a gallery image in Member Collections
From the album: Invertebrates
Same story as yesterday: My colleague and I wanted to visit a customer near Nincheng in Nei Mongol, China when I saw a drywall made out of grey flag stones. Looked very promising. I could pursuade my colleague (who was driving the car) to take a rest.... Unfortunately I could not tear down the wall - the owner of the drywall didn't like my idea and opposed. I guess the shale is from Daohugou region. Mecoptera non det. Female Possibly belonging to the Panorpidae Jiulongshan Formation Middle Jurassic Ningcheng Nei Mongol China- 1 comment
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From the album: Invertebrates
My colleague and I wanted to visit a customer near Nincheng in Nei Mongol, China when I saw a drywall made out of grey flag stones. Looked very promising. I could pursuade my colleague (who was driving the car) to take a rest.... Unfortunately I could not tear down the wall - the owner of the drywall didn't like my idea and opposed. I guess the shale came from Daohugou. Colorifuzia agenora Wei, Liang & Ren, 2013 Middle to Late Jurassic Jiulongshan Formation Daohugou Nei Mongol China-
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From the album: Invertebrates
Pseudopolycentropus daohugouensis ZHANG, 2005 Middle Jurassic Jiulongshan Formation Daohugou Nei Mongol China Lit.: Grimaldi, D.A., Junfeng, Z., Fraser, N.C., Rasnitsyn, A., 2005. Revision of the bizarre Mesozoic scorpionflies in the Pseudopolycentropodidae (Mecopteroidea). Insect Syst. Evol. 36, 443–458. Ren, D., Shih, C., Labandeira, C.C., 2010. New Jurassic pseudopolycentropodids from China (Insecta: Mecoptera). Acta Geol. Sinica 84, 22–30. Shih CK, Yang XG, Labandeira CC, Ren D (2011) A new long-proboscid genus of Pseudopolycentropodidae (Mecoptera) from the Middle Jurassic of China and its plant-host specializations. In: Shcherbakov DE, Engel MS, Sharkey MJ (Eds) Advances in the Systematics of Fossil and Modern Insects: Honouring Alexandr Rasnitsyn. ZooKeys 130: 281–297. doi: 10.3897/zookeys.130.1641-
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References: Grimaldi, D.A., Junfeng, Z., Fraser, N.C., Rasnitsyn, A., 2005. Revision of the bizarre Mesozoic scorpionflies in the Pseudopolycentropodidae (Mecopteroidea). Insect Syst. Evol. 36, 443–458. Ren, D., Shih, C., Labandeira, C.C., 2010. New Jurassic pseudopolycentropodids from China (Insecta: Mecoptera). Acta Geol. Sinica 84, 22–30. Shih CK, Yang XG, Labandeira CC, Ren D (2011) A new long-proboscid genus of Pseudopolycentropodidae (Mecoptera) from the Middle Jurassic of China and its plant-host specializations. In: Shcherbakov DE, Engel MS, Sharkey MJ (Eds) Advances in the Systematics of Fossil and Modern Insects: Honouring Alexandr Rasnitsyn. ZooKeys 130: 281–297. doi: 10.3897/zookeys.130.1641
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